Succulent

Succulent plants, such as this Aloe, store water in their fleshy leaves

Succulent plants, also known as succulents or fat plants, are water-retaining plants adapted to xerophilic climatic or soil conditions. Succulent plants store water in their leaves, stems and/or roots. The storage of water often gives succulent plants a more swollen or fleshy appearance than other plants, also known as succulence. In addition to succulence, succulent plants variously have other water-saving features. These may include:

• CAM metabolism to minimise water loss
• Absent, reduced, or cylindrical to spherical leaves
• reduction in the number of stomata
• stems, rather than leaves, as the main site of photosynthesis
• a compact, reduced, cushion-like, columnar or spherical growth form
• ribs enabling rapid increases in plant volume and decreasing surface area exposed to the sun
• waxy, hairy or spiny outer surface to reduce water loss via the creation of a humid microhabitat around the plant and a reduction in air movement near the surface of the plant.

Many succulents come from the dry areas of the tropics and subtropics, such as steppes, semi-desert and desert. High temperatures and low precipitation force plants to collect and store water in order to survive long dry periods. Succulents also occur as epiphytes, as such they have limited or no contact with the ground, and are dependent on their ability to store water. Succulents also occur as inhabitants of sea coasts, or salt pans which are exposed to high levels of dissolved minerals.

The best known succulents are cacti (family: Cactaceae). Virtually all cacti are succulents, but many succulents are not cacti.

Families and genera

Aizoaceae:Lithops julii, leaf succulent

Apocynaceae:Pachypodium lealii, stem succulent

Asphodelaceae:Haworthia arachnoidea, leaf succulent

File:Adansonia gregorii.jpg

Bombaceae:Adansonia gregorii, stem succulent

Cactaceae:Rebutia muscula, stem succulent

Crassulaceae:Crassula ovata, stem and leaf succulent

Euphorbiaceae:Euphorbia obesa ssp. symmetrica, stem succulent

Moringaceae:Moringa ovalifolia, stem succulent

Nolinaceae:Beaucarnea recurvata, stem succulent

Ruscaceae:Dracaena draco, stem succulent

Plant families and genera in which succulent species occur are listed below.

• Agavaceae: Agave, Beschorneria, Chlorophytum, Furcraea, Hesperaloe, Hesperoyucca, Yucca
• Aizoaceae: Acrodon, Aloinopsis, Amoebophyllum, Amphibolia, Antegibbaeum, Apatesia, Aptenia, Arenifera, Argyroderma, Aridaria, Astridia, Bergeranthus, Berrisfordia, Braunsia, Brownanthus, Calamophyllum, Carpobrotus, Carruanthus, Caryotophora, Cephalophyllum, Cerochlamys, Chasmatophyllum, Cheiridopsis, Conicosia, Conophytum, Cylindrophyllum, Dactylopsis, Delosperma, Dicrocaulon, Dinteranthus, Dorotheanthus, Dracophilus, Drosanthemum, Eberlanzia, Ebracteola, Ectotropis, Enarganthe, Erepsia, Esterhuysenia, Eurystigma, Faucaria, Fenestraria, Frithia, Gibbaeum, Glottiphyllum, Halenbergia, Hereroa, Herrea, Herreanthus, Hydrodea, Hymenogyne, Imitaria, Jacobsenia, Juttadinteria, Kensitia, Khadia, Lampranthus, Lapidaria, Leipoldtia, Lithops, Machairophyllum, Malephora, Maughaniella, Mesembryanthemum, Mestoklama, Meyerophytum, Micropterum, Mimetophytum, Mitrophyllum, Monilaria, Muiria, Namaquanthus, Namibia, Nanathus, Nelia, Neohenricia, Neorhine, Nycteranthus, Octopoma, Odontophorus, Oophytum, Ophthalmophyllum, Orthopterum, Oscularia, Ottosonderia, Pherelobus, Platythyra, Pleiospilos, Polymita, Prenia, Psammophora, Psicaulon, Rabiea, Rhinephyllum, Rhombophyllum, Ruschia, Ruschianthemum, Ruschianthus, Saphesia, Sceletium, Schwantesia, Scopologena, Semnanthe, Skiatophytum, Smicrostigma, Sphalmanthus, Stayneria, Stomatium, Synaptophyllum, Titanopsis, Trichodiadema, Vanheerea, Vanzijlia, Wooleya, Zeuktophyllum
• Amaranthaceae: Arthraerva, Salicornia
• Amaryllidaceae: Boophane, Brunsvigia, Cyrtanthus, Haemanthus, Rauhia
• Anacardiaceae: Operculicaria, Pachycormus
• Apiaceae: Steganotaenia
• Apocynaceae: Adenium, Mandevilla, Pachypodium, Plumeria
  • subfamily Asclepiadoideae (syn. Asclepiadaceae): Absolmsia, Asclepias, Aspidoglossum, Aspidonepsis, Baynesia, Brachystelma, Caralluma, Ceropegia, Cibirhiza, Cynanchum, Dischidia, Dischidiopsis, Duvalia, Duvaliandra, Echidnopsis, Edithcolea, Fanninia, Fockea, Glossostelma, Hoodia, Hoya, Huernia, Huerniopsis, Ischnolepis, Larryleachia, Lavrania, Madangia, Marsdenia, Matelea, Micholitzia, Miraglossum, Notechidnopsis, Odontostelma, Ophionella, Orbea, Orbeanthus, Pachycarpus, Pectinaria, Petopentia, Piaranthus, Pseudolithos, Quaqua, Raphionacme, Rhytidocaulon, Riocreuxia, Sarcorrhiza, Sarcostemma, Schizoglossum, Schlechterella, Stapelia, Stapelianthus, Stapeliopsis, Stathmostelma, Stenostelma, Stomatostemma, Tavaresia, Trachycalymma, Tridentea, Tromotriche, White-sloanea, Xysmalobium
• Araceae: Zamioculcas
• Araliaceae: Cussonia
• Asparagaceae: Myrsiphyllum
• Asphodelaceae: Aloe, Astroloba, Bulbine, Chortolirion, Gasteria, Haworthia, Poellnitzia, Trachyandra
• Asteraceae: Baeriopsis, Coulterella, Crassocephalum, Didelta, Gynura, Osteospermum, Othonna, Polyachyrus, Pteronia, Senecio
• Balsaminaceae: Impatiens
• Basellaceae: Anredera, Basella
• Begoniaceae: Begonia
• Bombaceae: Adansonia, Cavanillesia, Ceiba, Pseudobombax
• Brassicaceae: Heliophila, Lepidium
• Bromeliaceae: Abromeitiella
• Burseraceae: Beiselia, Bursea, Commiphora
• Cactaceae: Acanthocalycium, Acanthocereus, Ariocarpus, Armatocereus, Arrojadoa, Arthrocereus, Astrophytum, Austrocactus, Aztekium, Bergerocactus, Blossfeldia, Brachycereus, Browningia, Brasilicereus, Calymmanthium, Carnegiea, Cephalocereus, Cephalocleistocactus, Cereus, Cintia, Cipocereus, Cleistocactus, Coleocephalocereus, Copiapoa, Corryocactus, Coryphantha, Dendrocereus, Denmoza, Discocactus, Disocactus, Echinocactus, Echinocereus, Echinopsis, Epiphyllum, Epithelantha, Eriosyce, Escobaria, Escontria, Espostoa, Espostoopsis, Eulychnia, Facheiroa, Ferocactus, Frailea, Geohintonia, Gymnocalycium, Haageocereus, Harrisia, Hatiora, Hylocereus, Jasminocereus, Lasiocereus, Leocereus, Lepismium, Leptocereus, Leuchtenbergia, Lophophora, Maihuenia, Malacocarpus, Mammillaria, Mammilloydia, Matucana, Melocactus, Micranthocereus, Mila, Monvillea, Myrtillocactus, Neobuxbaumia, Neolloydia, Neoraimondia, Neowerdermannia, Obregonia, Opuntia, Oreocereus, Oroya, Ortegocactus, Pachycereus, Parodia, Pediocactus, Pelecyphora, Peniocereus, Pereskia, Pereskiopsis, Pilosocereus, Polaskia, Praecereus, Pseudoacanthocereus, Pseudorhipsalis, Pterocactus, Pygmaeocereus, Quiabentia, Rauhocereus, Rebutia, Rhipsalis, Samaipaticereus, Schlumbergera, Sclerocactus, Selenicereus, Stenocactus, Stenocereus, Stephanocereus, Stetsonia, Strombocactus, Tacinga, Thelocactus, Turbinicarpus, Uebelmannia, Weberbauerocereus, Weberocereus, Yungasocereus
• Campanulaceae: Brighamia
• Capparidaceae: Maerua
• Caricaceae: Carica, Jacarathia
• Chenopodiaceae
• Cochlospermaceae
• Commelinaceae: Aneilema, Callisia, Cyanotis, Tradescantia, Tripogandra
• Convolvulaceae: Ipomea, Sictocardia, Turbina
• Crassulaceae: Adromischus, Aeonium, Afrovivella, Aichryson, Cotyledon, Crassula, Cremnophila, Cremnosedum, Dudleya, Echeveria, Graptopetalum, Hylotelephium, Hypagophytum, Kalanchoe, Lenophyllum, Meterostachys, Monanthes, Orostachys, Pachyphytum, Perrierosedum, Phedimus, Pistorinia, Prometheum, Pseudosedum, Rhodiola, Rosularia, Sedella, Sedum, Sempervivum, Sinocrassula, Thompsonella, Tylecodon, Umbilicus, Villadia
• Cucurbitaceae: Apodanthera, Brandegea, Cephalopentandra, Ceratosanthes, Citrullus, Coccinia, Corallocarpus, Cucumella, Cucumis, Cucurbita, Cyclantheropsis, Dendrosicyos, Doyera, Eureindra, Fevillea, Gerrandanthus, Gynostemma, Halosicyos, Ibervilla, Kedostris, Marah, Momordica, Neoalsomitra, Odosicyos, Parasicyos, Syrigia, Telfairia, Trochomeria, Trochomeriopsis, Tumamoca, Xerosicyos, Zehneria, Zygosicyos
• Didiereaceae: Alluaudia, Alluaudiopsis, Decaria, Didierea
• Dioscoreaceae: Dioscorea
• Doryanthaceae: Doryanthes
• Ericaceae: Sphyrospermum
• Eriospermaceae: Eriospermum
• Euphorbiaceae: Cnidoscolus, Euphorbia, Jatropha
• Fabaceae: Delonix, Dolichos, Erythrina, Neorautanenia, Pachyrhizus, Tylosema
• Fouquieriaceae: Fouquieria
• Geraniaceae: Monsonia, Pelargonium
• Gesneriaceae: Aeschynanthus, Alsobia, Chirita, Codonanthe, Columnea, Nematanthus, Sinningia, Streptocarpus
• Hyacinthaceae: Albuca, Bowiea, Dipcadi, Drimia, Hyacinthus, Lachenalia, Ledebouria, Litanthus, Massonia, Ornithogalum, Rhadamanthus, Rhodocodon, Schizobasis, Urginea, Whiteheadia
• Icacinaceae: Pyrenacantha
• Lamiaceae: Aeollanthus, Dauphinea, Perrierastrum, Plectranthus, Solenostemon, Tetradenia, Thorncroftia
• Lentibulariaceae
• Loasaceae: Schismocarpus
• Loranthaceae: Tapinanthus
• Melastomataceae: Medinilla
• Meliaceae: Entandrophragma
• Menispermaceae: Chasmanthera, Stephania, Tinospora
• Moraceae: Dorstenia, Ficus
• Moringaceae: Moringa
• Nolanaceae: Nolana
• Nolinaceae: Beaucarnea, Calibanus, Dasylirion, Nolina
• Orchidaceae subfamily Epidendroideae Phalaenopsis
• Oxalidaceae: Oxalis
• Passifloraceae: Adenia
• Pedaliaceae: Pterodiscus, Sesamothamnus, Uncarina
• Phyllanthaceae: Phyllanthus
• Phytolaccaceae: Phytolacca
• Piperaceae: Peperomia
• Portulacaceae: Amphipetalum, Anacampseros, Avonia, Calyptrotheca, Ceraria, Cistanthe, Dendroportulaca, Grahamia, Lewisia, Parakeelya, Portulaca, Portulacaria, Schreiteria, Talinella, Talinum
• Rubiaceae: Anthorrhiza, Hydnophythum, Hydrophylax, Myrmecodia, Myrmephythum, Phylohydrax, Squamellaria
• Ruscaceae: Cordyline, Dracaena, Sansevieria
• Sapindaceae: Erythrophysa
• Saxifragaceae
• Sterculiaceae: Brachychiton, Sterculia
• Urticaceae: Laportea, Obertia, Pilea, Sarcopilea
• Viscaceae: Viscum
• Vitaceae: Cissus, Cyphostemma
• Xanthorrhoeaceae: Xanthorrhoea
• Zygophyllaceae

For some families, most members are succulent; for example the Cactaceae, Agavaceae, Aizoaceae, and Crassulaceae.

The table below shows the number of succulent species found in some families:

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Family	Succulent #	Modified parts	Distribution
Agavaceae	300	Leaf	North and Central America
Cactaceae	1600	Stem (root, leaf)	The Americas
Crassulaceae	1300	Leaf (root)	Worldwide
Aizoaceae	2000	Leaf	Southern Africa
Apocynaceae	500	Stem	Africa, Arabia, India
Didiereaceae	11	Stem	Madagascar (endemic)
Euphorbiaceae	> 1000	Stem and/or leaf and/or root	Africa, Madagascar, Asia, the Americas, Europe, Australia
Asphodelaceae	500	Leaf	Africa, Madagascar
Portulacaceae	?	Leaf and stem	The Americas