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{{Taxobox
| color = lightgreen
| name = ''Toxicodendron''
| image = Toxicodendron radicans.jpg
| image_width = 240px
| image_caption = ''[[Toxicodendron radicans]]''
| regnum = [[Plant]]ae
| divisio = [[Flowering plant|Magnoliophyta]]
| classis = [[Dicotyledon|Magnoliopsida]]
| ordo = [[Sapindales]]
| familia = [[Anacardiaceae]]
| genus = '''''Toxicodendron'''''
| genus_authority = [[Gerrit Smith Miller|Miller]]
| subdivision_ranks = Species
| subdivision = See text
}}
'''''Toxicodendron''''' is a genus of woody [[tree]]s, [[shrub]]s and [[vine]]s in the [[Anacardiaceae]] or Sumac Family. All members of the genus produce the skin-irritating oil [[urushiol]], which can cause a severe allergic reaction; hence the scientific name which means "poison tree".
Members of this genus are sometimes included in the genus ''[[Rhus]]'', although doing so leads to a [[polyphyletic]] ''Rhus''. The ''Toxicodendron'' genus is monophyletic.<ref>{{cite web | url=http://etd.lsu.edu/docs/available/etd-04152004-101232/ | title=Molecular Systematics of the Cashew Family (Anacardiaceae) (PhD dissertation at Louisiana State University) | author=Pell, Susan Katherine | date=[[2004-02-18]]}}, page 89</ref>
They have pinnately compound, alternate leaves and whitish or grayish drupes. The best known member is [[Poison-ivy (plant)|poison-ivy]], practically ubiquitous throughout much of eastern [[North America]].
The plants are quite variable in appearance. The leaves may have smooth, toothed or lobed edges, and all three types of leaf edge may be present in a single plant. The plants grow as creeping vines, climbing vines, shrubs, or, in the case of Lacquer Tree and Poison Sumac, as trees. While leaves of Poison-ivy and poison-oaks usually have three leaflets, sometimes there are five or, occasionally, even seven leaflets. Leaves of Poison Sumac have 7-13 leaflets, and of Lacquer Tree, 7-19 leaflets.
The common names come from similar appearances to other species that are not closely related and to the allergic response to the urushiol. Poison-oak is not an [[oak]] (''Quercus'', family [[Fagaceae]]), but this common name comes from the leaves' resemblance to white oak (''[[Quercus alba]]'') leaves, while Poison-ivy is not an [[ivy]] (''Hedera'', family [[Araliaceae]]), but has a superficially similar growth form. Both Poison-oak and Poison-ivy are members of the [[sumac]] family, Anacardiaceae. Technically, the plants do not contain a [[poison]]; they contain a potent [[allergen]].
The berries of certain species native to Japan and China, such as ''T. vernicifluum'' ([[Lacquer Tree]]) and ''T. succedaneum'' ([[Wax Tree]]), are used to make japan wax.
==Avoidance, treatment, and safety==
''For specific information on prevention and treatment of ''Toxicodendron'' rashes, see '''[[Urushiol-induced contact dermatitis]]'''.''
==Species of ''Toxicodendron''==
*[[Western Poison-oak]] (''Toxicodendron diversilobum'' or ''Rhus diversiloba'') is found only on the Pacific Coast of [[North America]], ranging from southern [[Canada]] to [[Baja California (peninsula)|Baja California]]. It is extremely common in that region, where it is the predominant species of the genus. Extremely variable, it grows as a dense shrub in open sunlight, or as a climbing vine in shaded areas. Like poison-ivy, it reproduces by creeping rootstocks or by seeds. The leaves are divided into three leaflets, 35-100 mm long, with scalloped, toothed, or lobed edges. Californians learn to recognize it by the rhyme "leaves of three, let it be". The leaves may be red, yellow, green, or some combination of those colors, depending on various factors, such as the time of year.
*[[Asian Poison-ivy]] (''Toxicodendron orientale'' or ''Rhus orientale'') is very similar to the American Poison-ivy, and replaces it throughout east Asia (so similar that some texts treat it as just a variety of the American species).
*[[Potanin's Lacquer Tree]] or Chinese Varnish Tree (''Toxicodendron potaninii'' or ''Rhus potaninii'') from central China, is similar to ''T. vernicifluum'' but with (usually) fewer leaflets per leaf. Growing up to 20 m tall, like ''T. vernicifluum'' it is used for [[lacquer]] production. The leaves have 7-9 leaflets.
*[[Poison-ivy (plant)|Poison-ivy]] (''Toxicodendron radicans'' or ''Rhus radicans'') is extremely common in some areas of North America. In the United States it grows in all states except Alaska, Hawaii, and California, but it is much less common than Poison-oak in western North America. It also grows in Central America. Appearing as a creeping vine, a climbing vine, or a shrub, it reproduces both by creeping rootstocks and by seeds. The appearance varies. Leaves, arranged in an alternate pattern, usually in groups of three, are from 20 to 50 mm long, pointed at the tip, and may be toothed, smooth, or lobed, but never serrated. Leaves may be shiny or dull, and the color varies with the season. Vines grow almost straight up rather than wrapping around their support, and can grow to 8-10 m in height. In some cases, Poison-ivy may entirely engulf the supporting structure, and vines may extend outward like limbs, so that it appears to be a Poison-ivy "tree".
*[[Western Poison-ivy]] (''Toxicodendron rydbergii'' or ''Rhus rydbergii'') is found in northern parts of the eastern United States. It also exists in the western United States and Canada, but is much less common than Poison-oak. It may grow as a vine or a shrub. It was once considered a subspecies of Poison-ivy. It does sometimes hybridize with the climbing species. Western Poison-ivy is found in much of western and central United States and Canada, although not on the West Coast. In the eastern United States it is rarely found south of New England.
*[[Wax Tree]] (''Toxicodendron succedaneum'' or ''Rhus succedanea''), a native of Asia, although it has been planted elsewhere, most notably Australia and New Zealand. It is a large shrub or tree, up to 8 m tall, somewhat similar to a sumac tree. Because of its beautiful autumn foliage, it has been planted outside of Asia as an [[ornamental plant]], often by gardeners who were apparently unaware of the dangers of allergic reactions. It is now officially classified as a noxious weed in Australia and New Zealand. The fatty-acid methyl ester of the kernel oil meets all of the major [[biodiesel]] requirements in the USA (ASTM D 6751-02, ASTM PS 121-99), Germany (DIN V 51606) and European Union (EN 14214). <ref>http://dx.doi.org/10.1016/j.biombioe.2005.05.001</ref>
*[[Atlantic Poison-oak]] (''Toxicodendron pubescens'' or ''Rhus toxicarium'') grows mostly in sandy soils in eastern parts of the United States. Growing as a shrub, its leaves are in groups of three. Leaves are typically rounded or lobed, and are densely haired. Although it is often confused with the more common poison-ivy, even in the scientific literature<ref>{{cite web | author=Sullivan, Janet | url=http://www.fs.fed.us/database/feis/plants/shrub/toxtox/all.html | title=Toxicodendron toxicarium | work=Fire Effects Information System | publisher=U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory | date=1994 | }}</ref>, Atlantic Poison-oak has small clumps of hair on the veins on the underside of the leaves, while Poison-ivy does not.
*[[Lacquer Tree]] or Varnish Tree (''Toxicodendron vernicifluum'' or ''Rhus verniciflua'') grows in Asia, especially China and Japan. Growing up to 20 m tall, its sap produces an extremely durable [[lacquer]]. The leaves have 7-19 leaflets (most often 11-13). The sap contains the allergenic oil, urushiol. Urushiol gets its name from this species which in [[Japanese language|Japanese]] is called ''Urushi''. Other names for this species include Japanese lacquer tree, Japanese Varnish Tree and Japanese Sumac (Note: the term "varnish tree" is also occasionally applied to the [[Candlenut]], ''Aleurites moluccana'', a southeast Asian tree unrelated to ''Toxicodendron'').
*[[Poison Sumac]] (''Toxicodendron vernix'' or ''Rhus vernix'') is a tall shrub or a small tree, from 2-7 m tall. It is found in swampy, open areas and reproduces by seeds. The leaves have between 7-13 untoothed leaflets, in a [[pinnate|feather-compound]] arrangement.<ref>{{cite book | title=A Field Guide to Eastern Trees | author=George A. Petrides | isbn=0-395-90455-2 | year=1998}}</ref> In terms of its potential to cause [[urushiol-induced contact dermatitis]], poison sumac is far more virulent than other ''Toxicodendron'' species, even more virulent than poison ivy and poison oak. According to some botanists, poison sumac (''Toxicodendron vernix'') is the most toxic plant species in the United States (Frankel, 1991).
==References==
<references />
==See also==
*[[Poison-ivy (plant)|Poison ivy]]
*[[Toxin]]
*[[Toxicodendron striatum]]
==External links==
*[http://www.pentrace.com/east/wajima/urushi.html Article about urushi lacquer made from the sap of the Urushi tree of Japan]
*[http://waynesword.palomar.edu/ww0802.htm Article about poison-oak]
*[http://herbarium.uvsc.edu/Virtual/search.asp?s=genus&p=1&n=14915&t=Toxicodendron UVSC Herbarium - Toxicodendron]
* Frankel, Edward, Ph.D. 1991. Poison Ivy, Poison Oak, Poison Sumac and Their Relatives; Pistachios, Mangoes and Cashews. The Boxwood Press. Pacific Grove, CA. (call #QK 495.A498 F73 1991). 98pages.
*[http://dx.doi.org/10.1016/j.biombioe.2005.05.001 Prospects and potential of fatty acid methyl esters of some non-traditional seed oils for use as biodiesel in India]
[[Category:Sapindales]]
[[Category:Poisonous_plants]]
[[Category:Dermatology]]
| color = lightgreen
| name = ''Toxicodendron''
| image = Toxicodendron radicans.jpg
| image_width = 240px
| image_caption = ''[[Toxicodendron radicans]]''
| regnum = [[Plant]]ae
| divisio = [[Flowering plant|Magnoliophyta]]
| classis = [[Dicotyledon|Magnoliopsida]]
| ordo = [[Sapindales]]
| familia = [[Anacardiaceae]]
| genus = '''''Toxicodendron'''''
| genus_authority = [[Gerrit Smith Miller|Miller]]
| subdivision_ranks = Species
| subdivision = See text
}}
'''''Toxicodendron''''' is a genus of woody [[tree]]s, [[shrub]]s and [[vine]]s in the [[Anacardiaceae]] or Sumac Family. All members of the genus produce the skin-irritating oil [[urushiol]], which can cause a severe allergic reaction; hence the scientific name which means "poison tree".
Members of this genus are sometimes included in the genus ''[[Rhus]]'', although doing so leads to a [[polyphyletic]] ''Rhus''. The ''Toxicodendron'' genus is monophyletic.<ref>{{cite web | url=http://etd.lsu.edu/docs/available/etd-04152004-101232/ | title=Molecular Systematics of the Cashew Family (Anacardiaceae) (PhD dissertation at Louisiana State University) | author=Pell, Susan Katherine | date=[[2004-02-18]]}}, page 89</ref>
They have pinnately compound, alternate leaves and whitish or grayish drupes. The best known member is [[Poison-ivy (plant)|poison-ivy]], practically ubiquitous throughout much of eastern [[North America]].
The plants are quite variable in appearance. The leaves may have smooth, toothed or lobed edges, and all three types of leaf edge may be present in a single plant. The plants grow as creeping vines, climbing vines, shrubs, or, in the case of Lacquer Tree and Poison Sumac, as trees. While leaves of Poison-ivy and poison-oaks usually have three leaflets, sometimes there are five or, occasionally, even seven leaflets. Leaves of Poison Sumac have 7-13 leaflets, and of Lacquer Tree, 7-19 leaflets.
The common names come from similar appearances to other species that are not closely related and to the allergic response to the urushiol. Poison-oak is not an [[oak]] (''Quercus'', family [[Fagaceae]]), but this common name comes from the leaves' resemblance to white oak (''[[Quercus alba]]'') leaves, while Poison-ivy is not an [[ivy]] (''Hedera'', family [[Araliaceae]]), but has a superficially similar growth form. Both Poison-oak and Poison-ivy are members of the [[sumac]] family, Anacardiaceae. Technically, the plants do not contain a [[poison]]; they contain a potent [[allergen]].
The berries of certain species native to Japan and China, such as ''T. vernicifluum'' ([[Lacquer Tree]]) and ''T. succedaneum'' ([[Wax Tree]]), are used to make japan wax.
==Avoidance, treatment, and safety==
''For specific information on prevention and treatment of ''Toxicodendron'' rashes, see '''[[Urushiol-induced contact dermatitis]]'''.''
==Species of ''Toxicodendron''==
*[[Western Poison-oak]] (''Toxicodendron diversilobum'' or ''Rhus diversiloba'') is found only on the Pacific Coast of [[North America]], ranging from southern [[Canada]] to [[Baja California (peninsula)|Baja California]]. It is extremely common in that region, where it is the predominant species of the genus. Extremely variable, it grows as a dense shrub in open sunlight, or as a climbing vine in shaded areas. Like poison-ivy, it reproduces by creeping rootstocks or by seeds. The leaves are divided into three leaflets, 35-100 mm long, with scalloped, toothed, or lobed edges. Californians learn to recognize it by the rhyme "leaves of three, let it be". The leaves may be red, yellow, green, or some combination of those colors, depending on various factors, such as the time of year.
*[[Asian Poison-ivy]] (''Toxicodendron orientale'' or ''Rhus orientale'') is very similar to the American Poison-ivy, and replaces it throughout east Asia (so similar that some texts treat it as just a variety of the American species).
*[[Potanin's Lacquer Tree]] or Chinese Varnish Tree (''Toxicodendron potaninii'' or ''Rhus potaninii'') from central China, is similar to ''T. vernicifluum'' but with (usually) fewer leaflets per leaf. Growing up to 20 m tall, like ''T. vernicifluum'' it is used for [[lacquer]] production. The leaves have 7-9 leaflets.
*[[Poison-ivy (plant)|Poison-ivy]] (''Toxicodendron radicans'' or ''Rhus radicans'') is extremely common in some areas of North America. In the United States it grows in all states except Alaska, Hawaii, and California, but it is much less common than Poison-oak in western North America. It also grows in Central America. Appearing as a creeping vine, a climbing vine, or a shrub, it reproduces both by creeping rootstocks and by seeds. The appearance varies. Leaves, arranged in an alternate pattern, usually in groups of three, are from 20 to 50 mm long, pointed at the tip, and may be toothed, smooth, or lobed, but never serrated. Leaves may be shiny or dull, and the color varies with the season. Vines grow almost straight up rather than wrapping around their support, and can grow to 8-10 m in height. In some cases, Poison-ivy may entirely engulf the supporting structure, and vines may extend outward like limbs, so that it appears to be a Poison-ivy "tree".
*[[Western Poison-ivy]] (''Toxicodendron rydbergii'' or ''Rhus rydbergii'') is found in northern parts of the eastern United States. It also exists in the western United States and Canada, but is much less common than Poison-oak. It may grow as a vine or a shrub. It was once considered a subspecies of Poison-ivy. It does sometimes hybridize with the climbing species. Western Poison-ivy is found in much of western and central United States and Canada, although not on the West Coast. In the eastern United States it is rarely found south of New England.
*[[Wax Tree]] (''Toxicodendron succedaneum'' or ''Rhus succedanea''), a native of Asia, although it has been planted elsewhere, most notably Australia and New Zealand. It is a large shrub or tree, up to 8 m tall, somewhat similar to a sumac tree. Because of its beautiful autumn foliage, it has been planted outside of Asia as an [[ornamental plant]], often by gardeners who were apparently unaware of the dangers of allergic reactions. It is now officially classified as a noxious weed in Australia and New Zealand. The fatty-acid methyl ester of the kernel oil meets all of the major [[biodiesel]] requirements in the USA (ASTM D 6751-02, ASTM PS 121-99), Germany (DIN V 51606) and European Union (EN 14214). <ref>http://dx.doi.org/10.1016/j.biombioe.2005.05.001</ref>
*[[Atlantic Poison-oak]] (''Toxicodendron pubescens'' or ''Rhus toxicarium'') grows mostly in sandy soils in eastern parts of the United States. Growing as a shrub, its leaves are in groups of three. Leaves are typically rounded or lobed, and are densely haired. Although it is often confused with the more common poison-ivy, even in the scientific literature<ref>{{cite web | author=Sullivan, Janet | url=http://www.fs.fed.us/database/feis/plants/shrub/toxtox/all.html | title=Toxicodendron toxicarium | work=Fire Effects Information System | publisher=U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory | date=1994 | }}</ref>, Atlantic Poison-oak has small clumps of hair on the veins on the underside of the leaves, while Poison-ivy does not.
*[[Lacquer Tree]] or Varnish Tree (''Toxicodendron vernicifluum'' or ''Rhus verniciflua'') grows in Asia, especially China and Japan. Growing up to 20 m tall, its sap produces an extremely durable [[lacquer]]. The leaves have 7-19 leaflets (most often 11-13). The sap contains the allergenic oil, urushiol. Urushiol gets its name from this species which in [[Japanese language|Japanese]] is called ''Urushi''. Other names for this species include Japanese lacquer tree, Japanese Varnish Tree and Japanese Sumac (Note: the term "varnish tree" is also occasionally applied to the [[Candlenut]], ''Aleurites moluccana'', a southeast Asian tree unrelated to ''Toxicodendron'').
*[[Poison Sumac]] (''Toxicodendron vernix'' or ''Rhus vernix'') is a tall shrub or a small tree, from 2-7 m tall. It is found in swampy, open areas and reproduces by seeds. The leaves have between 7-13 untoothed leaflets, in a [[pinnate|feather-compound]] arrangement.<ref>{{cite book | title=A Field Guide to Eastern Trees | author=George A. Petrides | isbn=0-395-90455-2 | year=1998}}</ref> In terms of its potential to cause [[urushiol-induced contact dermatitis]], poison sumac is far more virulent than other ''Toxicodendron'' species, even more virulent than poison ivy and poison oak. According to some botanists, poison sumac (''Toxicodendron vernix'') is the most toxic plant species in the United States (Frankel, 1991).
==References==
<references />
==See also==
*[[Poison-ivy (plant)|Poison ivy]]
*[[Toxin]]
*[[Toxicodendron striatum]]
==External links==
*[http://www.pentrace.com/east/wajima/urushi.html Article about urushi lacquer made from the sap of the Urushi tree of Japan]
*[http://waynesword.palomar.edu/ww0802.htm Article about poison-oak]
*[http://herbarium.uvsc.edu/Virtual/search.asp?s=genus&p=1&n=14915&t=Toxicodendron UVSC Herbarium - Toxicodendron]
* Frankel, Edward, Ph.D. 1991. Poison Ivy, Poison Oak, Poison Sumac and Their Relatives; Pistachios, Mangoes and Cashews. The Boxwood Press. Pacific Grove, CA. (call #QK 495.A498 F73 1991). 98pages.
*[http://dx.doi.org/10.1016/j.biombioe.2005.05.001 Prospects and potential of fatty acid methyl esters of some non-traditional seed oils for use as biodiesel in India]
[[Category:Sapindales]]
[[Category:Poisonous_plants]]
[[Category:Dermatology]]