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{{Taxobox
| color = lightgreen
| name = ''Taxus baccata''
| status = LR/lc
| image = TXbaccata.jpg
| image_width = 240px
| image_caption = ''Taxus baccata'' (European Yew) shoot with mature and immature cones
| regnum = [[Plant]]ae
| divisio = [[Pinophyta]]
| classis = [[Pinophyta|Pinopsida]]
| ordo = [[Pinales]]
| familia = [[Taxaceae]]
| genus = ''[[Taxus]]''
| species = '''''T. baccata'''''
| binomial = ''Taxus baccata''
| binomial_authority = [[Carolus Linnaeus|L.]]
}}

'''''Taxus baccata''''' is a [[Pinophyta|conifer]] native to western, central and southern [[Europe]], northwest [[Africa]], northern [[Iran]] and southwest [[Asia]].<ref name=rushforth>Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins ISBN 0-00-220013-9.</ref> Originally and still widely known in English as just '''Yew''', the later discovery of other very similar related species has led to qualification as '''European Yew''', '''Common Yew''' or '''English Yew''' where detail of which species of ''[[Taxus]]'' is required.

==Description==
It is a small to medium-sized [[evergreen]] [[tree]], growing 10-20 m (exceptionally up to 28 m) tall, with a trunk up to 2 m (exceptionally 4 m) diameter. The [[bark]] is thin, scaly brown, coming off in small flakes aligned with the stem. The [[leaf|leaves]] are [[lanceolate]], flat, dark green, 1-4 cm long and 2-3 mm broad, arranged spirally on the stem, but with the leaf bases twisted to align the leaves in two flat rows either side of the stem, except on erect leading shoots where the spiral arrangement is more obvious.<ref name=rushforth/><ref name=afm2>Mitchell, A. F. (1972). ''Conifers in the British Isles''. Forestry Commission Booklet 33.</ref>

[[Image:YewSeed.jpg|left|thumb|European Yew cone (detail)]]
The [[conifer cone|seed cones]] are highly modified, each cone containing a single [[seed]] 4-7 mm long partly surrounded by a modified scale which develops into a soft, bright red [[berry]]-like structure called an [[aril]], 8-15 mm long and wide and open at the end. The arils are mature 6-9 months after pollination, and with the seed contained are eaten by [[thrush (bird)|thrushes]], [[waxwing]]s and other [[bird]]s, which disperse the hard seeds undamaged in their droppings; maturation of the arils is spread over 2-3 months, increasing the chances of successful seed dispersal. The male cones are globose, 3-6 mm diameter, and shed their [[pollen]] in early spring. It is mostly [[Plant sexuality|dioecious]], but occasional individuals can be variably [[Plant sexuality|monoecious]], or change sex with time.<ref name=rushforth/><ref name=afm2/><ref name=dj>Dallimore, W., & Jackson, A. B. (1966). ''A Handbook of Coniferae and Ginkgoaceae'' 4th ed. Arnold.</ref>

It is relatively slow growing, but can be very long-lived, with the maximum recorded trunk diameter of 4 m probably only being reached in about 2,000 years. The potential age of yews is impossible to determine accurately and is subject to much dispute. There is rarely any wood as old as the entire tree, while the boughs themselves often hollow with age, making [[dendrochronology|ring counts]] impossible. There are unconfirmed claims as high as 5,000-9,500 years,<ref name=lewington>Lewington, A., & Parker, E. (1999). ''Ancient Trees: Trees that Live for a Thousand Years''. London: Collins & Brown Ltd. ISBN 1-85585-704-9</ref> but other evidence based on growth rates and archaeological work of surrounding structures suggests the oldest trees (such as the [[Fortingall Yew]] in [[Perthshire]], [[Scotland]]) are more likely to be in the range of 2,000 years.<ref name=harte>Harte, J. (1996). How old is that old yew? ''At the Edge'' 4: 1-9. Available [http://www.indigogroup.co.uk/edge/oldyews.htm online].</ref><ref name=kinmonth>Kinmonth, F. (2006). Ageing the yew - no core, no curve? ''International Dendrology Society Yearbook'' 2005: 41-46.</ref> Even with this lower estimate, ''Taxus baccata'' is the longest living plant in Europe.

All parts of the tree are [[toxic]], except the bright red aril surrounding the seed, enabling ingestion and dispersal by birds. The major toxin is the alkaloid [[taxine]]. The foliage remains toxic even when wilted or dried. [[Horse]]s have the lowest tolerance, with a lethal dose of 200–400 mg/kg body weight, but [[cattle]], [[pig]]s, and other livestock are only slightly less vulnerable.<ref name=>Tiwary, A. K., Puschner, B., Kinde, H., & Tor, E. R. (2005). Diagnosis of Taxus (Yew) poisoning in a horse. ''J. Vet. Diagn. Invest''. 17: 252–255.</ref> Symptoms include staggering gait, muscle tremors, convulsions, collapse, difficulty breathing, and eventually heart failure. However, death occurs so rapidly that many times the symptoms are missed. The tree should be fenced off or removed from pasture land to prevent grazing animals from consuming it.

==Etymology==
The word ''yew'' is from [[Proto-Germanic]] ''*īwa-'', possibly originally a loanword from [[Gaulish language|Gaulish]] ''ivos'', compare [[Irish language|Irish]] ''[[iodhadh|ēo]]'', [[Welsh language|Welsh]] ''ywen'', French ''[[:fr:If (Botanique)|if]]''; see [[Eihwaz]] for a discussion). ''Baccata'' is Latin for ''bearing berries''.

==Uses and traditions==
In the ancient [[Celtic mythology|Celtic]] world, the yew tree (''*eburos'') had extraordinary importance; a passage by [[Julius Caesar|Caesar]] narrates that Catuvolcus, chief of the [[Eburones]], literally "farmers of the yew", poisoned himself with yew rather than submit to Rome (''Gallic Wars'' 6: 31). Similarly, [[Florus]] notes that when the [[Cantabri]]ans were under siege by the legate Gaius Furnius in [[22 BC]], most of them took their lives either by the sword or by fire or by a poison extracted ''ex arboribus taxeis'', that is, from the yew tree (2: 33, 50-51). In a similar way, [[Orosius]] notes that when the [[Galician_People|Gallaecians]] were besieged at ''Mons Medullius'', they preferred to die by their own swords or by the yew tree poison rather than surrender (6, 21, 1.).

[[Image:English Yew 600.jpg|left|thumb|An Irish Yew (''Taxus baccata'' 'Fastigiata') planted at [[Kenilworth Castle]]]]
The yew is often found in [[church]]yards from [[England]] and [[Ireland]] to [[Galicia]]; some of these trees are exceptionally large (over 3 m diameter) and may be up to 2,000 years old, though few if any predate the churches they are beside.<ref name=harte/><ref name=kinmonth/> Many believe that the enormous sacred evergreen at the [[Temple at Uppsala]] was a yew{{Fact|date=October 2007}}. The Christian church commonly found it expedient to take over these existing sacred sites for churches. It is sometimes suggested that these were planted as a symbol of long life or trees of death. An explanation that the yews were planted to discourage farmers and drovers from letting their animals wander into the burial grounds, with the poisonous foliage being the disincentive, may be intentionally prosaic.

Yew is also associated with [[Wales]] and [[England]] because of the [[longbow]], an early weapon of war developed in northern Europe, and as the [[English longbow]] the basis for a mediaeval tactical system. Yew is the wood of choice for longbow making; the bows are constructed so that the [[Wood#Heartwood and sapwood|heartwood]] of yew is on the inside of the bow while the [[Wood#Heartwood and sapwood|sapwood]] is on the outside. This takes advantage of the natural properties of yew wood since the heartwood is able to withstand [[physical compression|compression]] while the sapwood is [[elastic]] and allows the bow to stretch. Both tend to return to their original straightness when the arrow is released. Much yew is knotty and twisted, so unsuitable for bowmaking; most trunks do not give good staves and even in a good trunk much wood has to be discarded.

The trade of yew wood to England for longbows was such that it depleted the stocks of good-quality, mature yew over a huge area. The first documented import of yew bowstaves to England was in [[1294]]. In [[1350]] there was a serious shortage, and [[Henry IV of England]] ordered his royal bowyer to enter private land and cut yew and other woods. In [[1470]] compulsory [[archery]] practice was renewed, and hazel, ash, and laburnum were specifically allowed for practice bows. Supplies still proved insufficient, until by the [[Statute of Westminster]] in 1472, every ship coming to an English port had to bring four bowstaves for every tun. [[Richard III of England]] increased this to ten for every tun. This stimulated a vast network of extraction and supply, which formed part of royal monopolies in southern Germany and Austria. In 1483, the price of bowstaves rose from two to eight [[pound sterling|pounds]] per hundred, and in 1510 the [[Venice|Venetians]] would only sell a hundred for sixteen pounds. In 1507 the [[Holy Roman Emperor]] asked the [[Rulers of Bavaria|Duke of Bavaria]] to stop cutting yew, but the trade was profitable, and in 1532 the royal monopoly was granted for the usual quantity "if there are that many". In 1562, the Bavarian government sent a long plea to the Holy Roman Emperor asking him to stop the cutting of yew, and outlining the damage done to the forests by its selective extraction, which broke the canopy and allowed wind to destroy neighbouring trees. In 1568, despite a request from Saxony, no royal monopoly was granted because there was no yew to cut, and the next year Bavaria and [[Austria]] similarly failed to produce enough yew to justify a royal monopoly. Forestry records in this area in the 1600s do not mention yew, and it seems that no mature trees were to be had. The English tried to obtain supplies from the Baltic, but at this period bows were being replaced by [[Gunpowder warfare|guns]] in any case.<ref>''Yew: A History''. Hageneder F. Sutton Publishing, 2007. ISBN 978-0750945974.</ref>.

[[Image:English Yew close 250.jpg|right|thumb|Foliage of Irish Yew; note the leaves spreading all round the erect shoots]]
Yews are widely used in landscaping and ornamental [[horticulture]]. Well over 200 [[cultivar]]s of ''Taxus baccata'' have been named. The most popular of these are the "Irish Yew" (''Taxus baccata'' 'Fastigiata'), a fastigiate cultivar of the European Yew selected from two trees found growing in Ireland, and the several cultivars with yellow leaves, collectively known as "Golden Yew". A special use of the yew is for [[topiary]] garden sculpture, a use not uncommon for many of the more elaborate gardens of [[England]] and [[Scotland]].<ref name=afm2/><ref name=dj/>

The precursors of [[chemotherapy]] drug [[Paclitaxel]] can be derived from the leaves of European Yew, which is a more renewable source than the bark of the endangered Pacific Yew (''[[Taxus brevifolia]]''). This ended a point of conflict in the early 1990s; many environmentalists, including Al Gore, had opposed the harvesting of paclitaxel for cancer treatments. [[Docetaxel]] (another [[taxane]]) can then be obtained by semi-synthetic conversion from the precursors.

==Literary References==
*In [[Shakesphere]]'s [[Titus Andronicus]], Act 2 Scene 3, Tamora the Goth queen exclaims: "No sooner had they told this hellish tale\ But straight they told me they would bind me here\ Unto the body of a dismal yew"
*In the Irish myth "The Love of Chu Chulainn and Fand", the warrior and the goddess meet beneath a yew tree's head at every quarter moon.
*[[John Keats]] refers to the yew in his "Ode on Melancholy", writing, "Make not your rosary of yew-berries, / Nor let the beetle, nor the death moth be / Your mournful Psyche, nor the downy owl / A partner in your sorrow's mysteries..." (lines 5-8).
*In Alfred, Lord Tennyson's poem "In Memoriam: A.H.H." the yew above Arthur Hallam's grave is addressed: "Old yew, which graspest at the stones/ That name the underlying dead,/ Thy fibres net the dreamless head,/ Thy roots are wrapped about the bones" (II, ln. 1-4).
*A Yew tree is featured prominently in [[William Wordsworth]]'s poems "Lines Left Upon a Seat in a Yew Tree" and "Yew-trees".
*In [[Alexandre Dumas, père|Alexandre Dumas]]'s novel, [[The Count of Monte Cristo]], [[Edmond Dantès]] is imprisoned in the [[Château d'If]], which literally translates to "Castle of the Yew" ([[If (island)|If]] is a small island in France, and the name may or may not derive from the word which means ''yew'').
*George Bernard Shaw's ''Mrs. Warren's Profession'' uses a yew tree in the yard of Reverend Samuel Gardner.
*In Section V of Little Gidding from T.S. Eliot's Four Quartets (the last section of the poem), Eliot claims: "The moment of the rose and the moment of the yew-tree/ Are of equal duration". In his poem, "Ash-Wednesday", he mentions the yew five times: "The silent sister veiled in white and blue/ Between the yews, behind the garden god, / Whose flute is breathless, bent her head and signed but/ spoke no word" (IV); "Till the wind shake a thousand whispers from the yew" (IV); "Will the veiled sister between the slender/ Yew trees pray for those who offend her" (V); "But when the voices shaken from the yew-tree drift away/ Let the other yew be shaken and reply" (VI).
*In [[J. R. R. Tolkien]]'s ''[[The Silmarillion]]'', [[Beleg Strongbow]] uses a bow made of yew. In ''[[The Hobbit]]'', the eagle king complains of the men of Wilderland using bows made of yew to shoot at his people.
*The murderer in [[Agatha Christie]]'s mystery ''[[A Pocket Full of Rye]]'' uses taxine ([[taxol]]), a poison derived from yew, to kill the victim. The victim lives at Yewtree Lodge.
*In [[Ursula LeGuin]]'s [[Earthsea]] series, both the wizard Ged and the Master Summoner carry staves of yew.
*In [[J. K. Rowling]]'s [[Harry Potter]] series, [[Voldemort]] uses a wand made of yew.
*The Yew is the subject of Swedish author [[Gunnar D Hansson]]'s "lyrical monography" ''[[Idegransöarna]]'' (''The Yew-tree Islands'', 1994, untranslated to English). Hansson explores the yew in its uses (medicinal, lyrical, in place-names, etc) and its historical meaning. He speculates about the yew, and weaves a tale of prose poems, essays and lyrics, about the yew; the book takes the reader close to the yew in its relation to Hittites, Vikings, medicine, Robin Hood, Christmas, heathendom, etymology and mythology.
*The [[Great Chain of Being]], which proposes a strict, hierarchical order for the beings (divine entities, animals, and plants) in the universe, designates the yew as the lowest form of tree among plants.

==References==
{{Commons|Taxus baccata}}
{{Reflist}}
* Chetan, A. and Brueton, D. (1994) ''The Sacred Yew'', London: Arkana, ISBN 0-14-019476-2
* Conifer Specialist Group (1998) ''Taxus baccata'', In: IUCN 2006/UCN Red List of Threatened Species, [http://www.iucnredlist.org/search/details.php/42546/summ WWW page] (Accessed 3 February 2007)
* Hartzell, H. (1991) ''The yew tree: a thousand whispers: biography of a species'', Eugene: Hulogosi, ISBN 0-938493-14-0
* Simón, F. M. (2005) Religion and Religious Practices of the Ancient Celts of the Iberian Peninsula, ''[http://www.uwm.edu/Dept/celtic/ekeltoi/volumes/vol6/6_6/marco_simon_6_6.html e-Keltoi]'', v. 6, p. 287-345, ISSN 1540-4889 online

==External links==
* [http://www.druidnetwork.org/articles/theyew.html Notes on the Yew] Druid Network

[[Category:Taxaceae]]
[[Category:Medicinal plants]]
[[Category:Poisonous plants]]
[[Category:Plants poisonous to equines]]
[[Category:Flora of Europe]]
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