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Crataegus (ancient Greek name, derived from kratos, strength, on account of the hardiness of the wood). Rosaceae, subfam. Pomeae. Crategus. Hawthorn. Woody plants grown for their handsome foliage, attractive flowers and decorative fruit which, in a few species, is edible, and also for their picturesque habit: very valuable for ornament.

Crataegus (ancient Greek name, derived from kratos, strength, on account of the hardiness of the wood). Rosaceae, subfam. Pomeae. Crategus. Hawthorn. Woody plants grown for their handsome foliage, attractive flowers and decorative fruit which, in a few species, is edible, and also for their picturesque habit: very valuable for ornament.

Shrubs or small trees, usually spiny: Lvs. alternate, deciduous, stipulate, serrate, often lobed or pinnatifid: fls. white, in some varieties red, in corymbs, rarely solitary; petals and calyx-lobes 5; stamens 5-25, usually 10 or 20; styles 1-5: fr. a drupe-like pome, with 1-5 1-seeded bony stones.—A large genus, widely distributed in the temperate regions of the northern hemisphere, most abundant in N. Amer., where between 800 and 900 species have been described, while from the Old World only about 60 species are known. There exists no recent monograph of the genus; a systematic enumeration of the arborescent American species will be found in Sargent, "Manual of the Trees of North America," pp. 363-504; of the species of the southern states in Small, "Flora of the Southeastern United States," pp. 532-569; and of the species of the northeastern states in Gray's Manual, ed. 7, p. 460- 79, and in Britton and Brown, 111. Flor. (ed. 2) 2:294-321; for the species cult, in European gardens, see Lange, "Revisio Specierum Generis Crataegi" (1897), quoted below as Lange.

Shrubs or small trees, usually spiny: Lvs. alternate, deciduous, stipulate, serrate, often lobed or pinnatifid: fls. white, in some varieties red, in corymbs, rarely solitary; petals and calyx-lobes 5; stamens 5-25, usually 10 or 20; styles 1-5: fr. a drupe-like pome, with 1-5 1-seeded bony stones.—A large genus, widely distributed in the temperate regions of the northern hemisphere, most abundant in N. Amer., where between 800 and 900 species have been described, while from the Old World only about 60 species are known. There exists no recent monograph of the genus; a systematic enumeration of the arborescent American species will be found in Sargent, "Manual of the Trees of North America," pp. 363-504; of the species of the southern states in Small, "Flora of the Southeastern United States," pp. 532-569; and of the species of the northeastern states in Gray's Manual, ed. 7, p. 460- 79, and in Britton and Brown, 111. Flor. (ed. 2) 2:294-321; for the species cult, in European gardens, see Lange, "Revisio Specierum Generis Crataegi" (1897), quoted below as Lange.

The hawthorns are hardy ornamental shrubs and trees, mostly of dense and low growth, with handsome foliage, turning, in most species, to a brilliant coloring in the fall. Almost all have attractive white flowers, pink or crimson in some varieties of C. Oxpacantha and C. monogyna. Most of the species have very decorative  fruit which in C. Phaenopyrum, C. nitida, C. viridis, C. fecunda, C. pruinosa, C. Carrierei, C. persistens, C. Oxyacantha, C. monogyna and others persist on the trees until late into the winter, while some species, as C. Arnoldiana, ripen their large fruits, which soon drop, in August; also C. dahurica, C. sanguinea and the black- fruited C. nigra ripen about the same time, and C. submollis only a little later, but the earliest of all is the southern C. aestivalis, which ripens its fruits in May. This and the blue-fruited C. brachyacantha are among the most decorative hawthorns for the southern states. The fruit of C. aestivalis, and that of C. mexicana is made into preserves and jellies; also the fruits of the Molles group are suited for jelly-making, and in South Carolina an excellent jelly similar in quality and taste to Guava jelly is made from the fruits of some species of the Flavae group. In Europe, C, monogyna and C. Oxyacantha are counted among the best hedge plants; also many American species like C. Phaenopyrum, C. Crus-galli and possibly] C. macracantha, C. intricate, C. pastorum, C. rotundifolia, may be used for hedges, but they are stronger growers and cannot be pruned so closely as the European species. The hawthorns grow well in exposed positions and as a rule do not like much shade; they are not particular as to the soil, but grow best in limestone soil, also in a rich, loamy, somewhat moist one, and even in strong clay. Propagated by seeds, sown in fall or stratified; before stratifying, most of the pulp may be removed by laying the fruits in shallow piles and allowing them to decay. Then they are mixed with sand or sifted soil and buried in the ground or kept in boxes in a cool cellar. The young plants should not be allowed to remain over one year in the seed-beds, as they form long tap-roots and are then difficult to transplant. Varieties and rarer kinds are easily budded or grafted on seedling stock of C. Oxyacantha, or other common strong-growing species.  

The hawthorns are hardy ornamental shrubs and trees, mostly of dense and low growth, with handsome foliage, turning, in most species, to a brilliant coloring in the fall. Almost all have attractive white flowers, pink or crimson in some varieties of C. Oxpacantha and C. monogyna. Most of the species have very decorative  fruit which in C. Phaenopyrum, C. nitida, C. viridis, C. fecunda, C. pruinosa, C. Carrierei, C. persistens, C. Oxyacantha, C. monogyna and others persist on the trees until late into the winter, while some species, as C. Arnoldiana, ripen their large fruits, which soon drop, in August; also C. dahurica, C. sanguinea and the black- fruited C. nigra ripen about the same time, and C. submollis only a little later, but the earliest of all is the southern C. aestivalis, which ripens its fruits in May. This and the blue-fruited C. brachyacantha are among the most decorative hawthorns for the southern states. The fruit of C. aestivalis, and that of C. mexicana is made into preserves and jellies; also the fruits of the Molles group are suited for jelly-making, and in South Carolina an excellent jelly similar in quality and taste to Guava jelly is made from the fruits of some species of the Flavae group. In Europe, C, monogyna and C. Oxyacantha are counted among the best hedge plants; also many American species like C. Phaenopyrum, C. Crus-galli and possibly] C. macracantha, C. intricate, C. pastorum, C. rotundifolia, may be used for hedges, but they are stronger growers and cannot be pruned so closely as the European species. The hawthorns grow well in exposed positions and as a rule do not like much shade; they are not particular as to the soil, but grow best in limestone soil, also in a rich, loamy, somewhat moist one, and even in strong clay. Propagated by seeds, sown in fall or stratified; before stratifying, most of the pulp may be removed by laying the fruits in shallow piles and allowing them to decay. Then they are mixed with sand or sifted soil and buried in the ground or kept in boxes in a cool cellar. The young plants should not be allowed to remain over one year in the seed-beds, as they form long tap-roots and are then difficult to transplant. Varieties and rarer kinds are easily budded or grafted on seedling stock of C. Oxyacantha, or other common strong-growing species.  

The spines of crategus are modified branches (see Fig. 1096). The fruits are pomes (Fig. 1097), with structure similar to that of the apple. Alfred Rehder.

The spines of crategus are modified branches (see Fig. 1096). The fruits are pomes (Fig. 1097), with structure similar to that of the apple. Alfred Rehder.

The fruit can be sown broadcast in beds without any separation of the seeds, and heavily mulched until the spring of the second year, when the mulching should be removed. This method, however, is not considered good, and has been given up. The best way is to soak the fruits in water, and by maceration the seeds or nutlets are separated from the pulp, and the seeds will sink to the bottom of the tub or vessel. The seeds are Chen dried in the sun as they can then be handled easily. They are sown in "flats" of convenient size to handle, and piled up in the corner of the shade house and fitted tightly above each other to prevent mice getting at them. During this period of rest they must not be allowed to become dry. In the spring of the second year they are spread out to allow the seeds to germinate. Numbered zinc tags are nailed on the flats" and the corresponding numbers with the names of the species are recorded.

The fruit can be sown broadcast in beds without any separation of the seeds, and heavily mulched until the spring of the second year, when the mulching should be removed. This method, however, is not considered good, and has been given up. The best way is to soak the fruits in water, and by maceration the seeds or nutlets are separated from the pulp, and the seeds will sink to the bottom of the tub or vessel. The seeds are Chen dried in the sun as they can then be handled easily. They are sown in "flats" of convenient size to handle, and piled up in the corner of the shade house and fitted tightly above each other to prevent mice getting at them. During this period of rest they must not be allowed to become dry. In the spring of the second year they are spread out to allow the seeds to germinate. Numbered zinc tags are nailed on the flats" and the corresponding numbers with the names of the species are recorded.

The American hawthorns can be grafted readily on potted seedling stocks in the greenhouse in winter, any of the species in the Crus-galli group being good to use. They are grafted at the crown. This, however, is an unnecessary operation. All of the species of American hawthorns (and there are over 900 of them) come absolutely true from seed, and whilst they germinate slowly, they start to grow rapidly into plants of good stocky size from about two years after they germinate.

The American hawthorns can be grafted readily on potted seedling stocks in the greenhouse in winter, any of the species in the Crus-galli group being good to use. They are grafted at the crown. This, however, is an unnecessary operation. All of the species of American hawthorns (and there are over 900 of them) come absolutely true from seed, and whilst they germinate slowly, they start to grow rapidly into plants of good stocky size from about two years after they germinate.

Some of the species of American hawthorns have highly colored foliage in the fall. The species in the Pruinosue, Medioximae and Intricate groups have perhaps the most highly colored foliage. Notable examples are Crataegus opulens, C. diffusa, C. maineana, C. dissona, C. cognata, C. conspecta, C. promissa, C. exornata, C. perjucunda, C. foetida, and C. verecunda.

Some of the species of American hawthorns have highly colored foliage in the fall. The species in the Pruinosue, Medioximae and Intricate groups have perhaps the most highly colored foliage. Notable examples are Crataegus opulens, C. diffusa, C. maineana, C. dissona, C. cognata, C. conspecta, C. promissa, C. exornata, C. perjucunda, C. foetida, and C. verecunda.

The different species vary greatly in the time of ripening their fruits and in the period of duration. In many instances the fruit drops soon after ripening and in others hangs on for a long period. A selection of twenty-six species that would give a good fruit display from August until early or mid-winter, would be as follows: Crataegus matura, C. praecox, C. Arnoldiana, C. Dayana, C. Robesoniana, C. pedicellata, C. gloriosa, C. Ellwangeriana, C. lauta, C. submollis, C. champlainensis, C. arkansana, C. Dunbari, C. ferentaria, C. opulens, C. compta, C. gemmosa, C. livoniana, C. geneseensis, C. persimilis, C. maineana, C. Barryana, C. coccinioides, C. leiophylla, C. durobrivensis, and C. cordata. (See pp. 887-889 for some of these.) John Dunbar.

The different species vary greatly in the time of ripening their fruits and in the period of duration. In many instances the fruit drops soon after ripening and in others hangs on for a long period. A selection of twenty-six species that would give a good fruit display from August until early or mid-winter, would be as follows: Crataegus matura, C. praecox, C. Arnoldiana, C. Dayana, C. Robesoniana, C. pedicellata, C. gloriosa, C. Ellwangeriana, C. lauta, C. submollis, C. champlainensis, C. arkansana, C. Dunbari, C. ferentaria, C. opulens, C. compta, C. gemmosa, C. livoniana, C. geneseensis, C. persimilis, C. maineana, C. Barryana, C. coccinioides, C. leiophylla, C. durobrivensis, and C. cordata. (See pp. 887-889 for some of these.)

 

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The number of [[species]] in the genus depends on taxonomic interpretation, with numerous [[apomixis|apomictic microspecies]]; some botanists recognise a thousand or more species, while others reduce the number to 200 or fewer.

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Hawthorns provide food and shelter for many species of [[bird]]s and [[mammal]]s, and the [[flower]]s are important for many nectar-feeding [[insect]]s. Hawthorns are also used as food plants by the [[larva]]e of a large number of [[Lepidoptera]] species — see [[list of Lepidoptera which feed on Hawthorns]].

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Many species and hybrids are used as [[ornamental plant|ornamental]] and street trees. The Common Hawthorn is extensively used in Europe as a [[hedge (barrier)|hedge]] plant. Several [[cultivar]]s of the Midland Hawthorn ''C. laevigata'' have been selected for their pink or red flowers. Hawthorns are among the trees most recommended for water-conservation landscapes.

===Pests and diseases===

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==Selected species==

==Species==

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==Uses==

*C. altaica, Lange = C. Wattiana.{{SCH}}

===Culinary use===

The fruits of the species ''Crataegus pinnatifida'' (Chinese Hawthorn) are tart, bright red, and resemble small [[Malus|crabapple]] fruits. They are used to make many kinds of Chinese snacks, including [[haw flakes]] and ''[[Tang Hu Lu|tanghulu]]'' (糖葫芦). The fruits, which are called ''shānzhā'' ([[wiktionary:山|山]][[wiktionary:楂|楂]]) in Chinese, are also used to produce jams, jellies, juices, alcoholic beverages, and other drinks [http://www.itmonline.org/arts/crataegus.htm]. In [[South Korea]], a liquor called ''sansachun'' (산사춘) is made from the fruits.[http://www.soolsool.co.kr/English/product.htm]

*C. Baxteri, Sarg.=C. foetida.{{SCH}}

 

*C. beata, Sarg. Allied to C. pruinosa. Shrub, to 15 ft.: lvs. oblong-ovate, villous above while young: anthers dark maroon-color: fr. crimson, pruinose, ripens end of Sept. N. Y.{{SCH}}

The fruits of ''Crataegus pubescens'' are known in [[Mexico]] as ''tejocotes'' and are eaten raw, cooked, or in jam during the winter months. They are stuffed in the ''[[piñata]]s'' broken during the traditional pre-Christmas parties known as ''[[posadas]]''. They are also cooked with other fruits to prepare a Christmas punch. The mixture of ''tejocote'' paste, sugar, and chili powder produces a popular Mexican candy called ''rielitos'', which is manufactured by several brands.

*C. Beckwithae, Sarg. Allied to C. pastorum. Shrub or tree, to 18 ft.: lvs. ovate, usually truncate at the base, at maturity thin: calyx-lobes glandular-serrate: fr. subglobose, crimson, with 5 stones. N. Y.{{SCH}}

 

*C. bellula. Sarg. Related to C. pruinosa. Shrub, to 12 ft.; glabrous: Lvs. ovate, bluish green and lustrous above, 2-3 in. long: stamens 8-10: fr. dull crimson, bloomy, ¾ in. across, with usually 4 stones. Mich. 8.T.8. 1:56.{{SCH}}

In the southern United States fruits of three native species are collectively known as [[Mayhaw]]s and are made into jellies which are considered a great delicacy.

*C. berberifolia, Torr. A Gray. Related to C. Crus-galli. Lvs. obovate or obovate-oblong, obtuse, pubescent below, lustrous and nearly glabrous above, 1⅓-2 in. long: corymbs pubescent: anthers yellow: fr. orange with red cheek. La. S.S. 179.{{SCH}}

 

*C. Celsiana. Bosc. Shrub: lvs. pinnately lobed, slightly pubescent beneath: corymbs many-fld.: fr. ovoid, red. Origin unknown, probably hybrid of C. pentagyna.{{SCH}}

The leaves are edible and, if picked in the months of April and May, they are tender enough to be used in salads.<ref>Richard Mabey, ''Food for Free'', Collins, October 2001.</ref>

*C. champlainensis, Sarg. Allied to C. mollis. Tree, to 20 ft., spiny: Lvs. ovate, usually truncate at the base, lobed, glabrous above, pubescent on the veins below, 2-2½ in. long: corymbs villous, usually 4-5-fld.; stamens 10: fr. obovoid or ovoid, scarlet, ⅓ in. long, in Sept. Que. and Ont. to Vt. and N. Y. S.S. 13:669.{{SCH}}

 

*C. chloroearca, Maxim. Allied to C. sanguinea. Lvs. pinnately lobed. truncate at the base, with short lobes, glabrous at length: corymbs many-fld., nearly glabrous: fr. black, with green flesh. Japan.{{SCH}}

*C. cognata. Sarg. Closely related to C. pruinosa. Shrub, to 10 ft., spiny: Lvs. ovate, acute or acuminate, slightly lobed, dull bluish green, glabrous: corymbs 5-7-fld.; anthers yellow: fr. ovoid or pyriform, pruinose, dull crimson at maturity, over ⅓ in. long, in Oct. Mass.{{SCH}}

The dried fruits of ''Crataegus pinnatifida'' (called [[wikt:山|山]][[wikt:楂|楂]]  or ''shān zhā'' in Chinese) are used in [[naturopathic medicine]] and [[traditional Chinese medicine]], primarily as a digestive aid. A closely related species, ''Crataegus cuneata'' (Japanese Hawthorn, called ''sanzashi'' in Japanese) is used in a similar manner. Other species (especially ''Crataegus laevigata'') are used in Western herbal medicine, where the plant is believed to strengthen cardiovascular function [http://www.itmonline.org/arts/crataegus.htm]. In recent years, this use has been noted and adopted by Chinese herbalists as well [http://www.itmonline.org/arts/crataegus.htm]. Hawthorn is also used as an aid to lower blood pressure, and treat some heart related diseases.  

*C. compta, Sarg. (C. silvicola var. compta, Eggleston). Allied to C. pruinosa. Shrub, spiny, glabrous: Lvs. oblong-ovate, usually rounded at the base, slightly lobed, glabrous: corymbs many-fld.; stamens 7-10; anthers dark row: fr. obovoid, light cherry-red, ½ in. long, in Oct. W. N. Y. {{SCH}}

 

*C. conspicta, Sarg. Allied to C. pruinosa. Tree, to 20 ft., spiny: Lvs. broadly ovate, rounded or subcordate at the base, lobed, yellow-green, pubescent on the midrib below: corymbs 5-6-fld., slightly hairy, compact; anthem white: fr. subglobose, crimson, over ½ in. across, in Oct. Ont.{{SCH}}

;Clinical trials

Several clinical trials have assessed the ability of hawthorn to help improve exercise tolerance in people with NYHA class II cardiac insufficiency compared to placebo. One trial, at (300mg/day) for 4 to 8 weeks, found not difference from placebo. The second trial, including 78 subjects (600mg/day) for 8 weeks, found "significant improvement in exercise tolerance" and lower blood pressure and heart rate during exercise. The third trial, including 32 subjects (900mg/day) for 8 weeks, found improved exercise tolerance as well as a reduction in the "incidence and severity of symptoms such as dyspnea and fatigue decreased by approximately 50%" <ref>Bauman HHSFJL (2002). Hawthorn. ''Journal of Cardiovascular Nursing'' 16:1-8</ref>.

*C. cuneata, Sieb. & Zucc. Belongs to group Cuneata-. Shrub: young branchlets villous: Lvs. ahort-petioled, cuneate-obovate or cuneate-ohlong, crenate-serrate, glabrous and lustrous above, sparingly hairy below: corymbs villous, 3-7-fld.; stamens 20; anthers red: fr. red, with 5 stones, plain inside. China and Japan. L. I. 5.{{SCH}}

 

*C. cupulifera. Sarg. Allied to C. rotundifolia. Shrub, to 20 ft.: Lvs. obovate or rhombic, slightly lobed, lustrous and scabrate above: corymbs slightly villous; fls. cup-shaped; stamens 10, anthers pink: fr. scarlet, with 3-4 stones. N. Y.{{SCH}}

In the HERB-CHF (Hawthorn Extract Randomized Blinded Chronic HF Study) clinical trial, 120 patients took 450mg of hawthorn extract twice daily for 6 months in combination with standard therapy and a standardized exercise program. "No effects of hawthorn were seen on either quality-of-life endpoint (Tables 1 and 2), or when adjusted for LVEF" <ref>Aaronson K: HERB-CHF: Hawthorn Extract Randomized Blinded Chronic Heart Failure Trial. In, 2004</ref>.

 

*C. diffusa, Sarg. Allied to C. pruinosa. Intricately branched spiny shrub, to 15 ft., glabrous: Lvs. ovate, acuminate, rounded or cuneate at the base, dark bluish green and slightly scabrate above: corymbs 6-12-fld.; stamens 10: fr. subglobose, scarlet, less than ½ in. across , in Oct .W.N.Y.{{SCH}}

One study, consisting of 1011 patients taking one tablet (standardized to 84.3mg [[proanthocyanidin|procyanidin]]) twice daily for 24 weeks, found "improvements in clinical symptoms (such as fatigue, palpitations, and exercise dyspnea), performance and exercise tolerance test, and ejection fraction" <ref>Sweet JMRBV (2002). Hawthorn: Pharmacology and therapeutic uses. ''American Journal of Health-System Pharmacy'' 59: 417-422</ref>.

 

*C. dissona, Sarg. (C. pruinosa var. dissona, Eggleston). Allied to C. pruinosa. Slender spiny shrub, to 10 ft., glabrous: Lvs. ovate to rhombic, cuneate at the base, dark bluish green: corymbs 5-7-fld.; stamens 10; anthers purple: fr. subglobose. crimson, in Oct. Mass.{{SCH}}

The wood of some hawthorn species is very hard and resistant to rot. In rural North America it was prized for use as tool handles and fence posts.

 

*C. aurobrizensis, Sarg. Allied to C. coccinioides. Shrub, to 20 ft., spiny: Lvs. ovate, with. 3-4 pairs of short lobes, glabrous: corymbs glabrous; stamens 20: fr. bright red, in Oct. N. Y. S.T.S. 1:2. {{SCH}}

==Folklore==

*C. elliptica, Ait. (C. glandulosa, Moench. C. flava var. pubescens, Gray). Allied to C. flava. Lvs. broader, of firmer texture, more pubescent and glandular: fr. subglobose, red or yellow. Southern states. B.R. 22: 1890 (as C. spathulata). {{SCH}}

The custom of employing the flowering branches for decorative purposes on the 1st of May is of very early origin; but since the adoption of the [[Gregorian calendar]] in 1752, the tree has rarely been in full bloom in England before the second week of that month. In the Scottish Highlands the flowers may be seen as late as the middle of June. The hawthorn has been regarded as the emblem of hope, and its branches are stated to have been carried by the ancient Greeks in wedding processions, and to have been used by them to deck the altar of [[Hymenaios]]. The supposition that the tree was the source of [[Jesus]]'s crown of thorns gave rise doubtless to the tradition current among the French peasantry that it utters groans and cries on [[Good Friday]], and probably also to the old popular superstition in Great Britain and Ireland that ill-luck attended the uprooting of hawthorns. Branches of [[Glastonbury Abbey#Glastonbury Thorn|Glastonbury Thorn]], C. Oxyacantha, var. praecox, which flowers both in December and in spring, were formerly highly valued in England, on account of the legend that the tree was originally the staff of [[Joseph of Arimathea]].<ref>"Hawthorn" article in the 1911 Encyclopedia</ref>

*C. elliptico. Beadle, is C. senta, Beadle, a species allied to C. flava. S.S. 13:697. {{SCH}}

 

*C. elliptica, Mohr, is C. signata, Beadle, a species allied to C. Crus-galli. S.S. 13:644. {{SCH}}

In [[Celt|Celtic]] lore, the hawthorn plant was used commonly for rune inscriptions along with [[Taxaceae|Yew]] and [[Apple]]. It was once said to heal the broken heart.

*C. exornata, Sarg. Allied to C. pruinosa. Shrub, to 10 ft., spiny: Lvs. ovate, usually rounded at the base, slightly lobed, dark yellow-green and rough above: corymbs 5-6-fld.; stamens 7-10; anthers pink: fr. subglobose, scarlet, less than ½ in. across, in Sept. Ont. {{SCH}}

*C. Fazonii, Sarg. Allied to C. rotundifolia. Shrub, to 10 ft., spiny: lvs. broadly ovate, with 4-5 pairs of short lobes, nearly glabrous at maturity: corymbs villous; stamens 5-10: fr. dark crimson, in Sept. N. H. S.T.S. 1:60.{{SCH}}

*C. ferentaria, Sarg. Allied to C. macracantha. Intricately branched shrub, to 12 ft.. with stout spines: Lvs. rhombic or obovate, puberulous below on the veins: corymbs slightly villous; stamens 7-10; anthers white: fr. subglobose or ovoid, ⅓ in. long, scarlet, with usually 2 stones, ripening in Oct. W.N.Y.{{SCH}}

*C. flabellata, Spach (C. Grayana, Eggleston). Allied to C. pedicellata. Shrub, to 20 ft.: Lvs. ovate, with short acute lobes, at first sparingly hairy above and villous at the veins beneath: corymbs slightly villous; calyx-lobes sparingly glandular- serrate; stamens 20: fr. ovoid, crimson, with 3-5 stones, in Sept. Que.{{SCH}}

*C.faetida, Ashe (C. Baxteri, Sarg.). Allied to C. intricate. Intricately branched, spiny shrub, to 12 ft., glabrous: Lvs. ovate or oval: corymbs usually 5-6-fld.; calyx-lobes serrate: fr. subglobose. orange-red or red- brown, about ½ in. thick, with 3-4 stones, in Oct. Mass, to Ont. and Va.{{SCH}}

*C. Fontanesiana, Steud. Allied to C. Crus-galli. Lvs. elliptic or elliptic-lanceolate, almost glabrous, shining above: corymbs many-fld., pubescent: fr. red. Probably hybrid of C. Crus-galli.{{SCH}}

*C. Forbease, Sarg. Allied to C. pastorum. Shrub, to 15 ft.: Lvs. ovate to oval, cuneate or rounded at the base, slightly lobed: stamens 20; anthers dark rose-color: fr. globose or ovoid, scarlet, with thin and juicy flesh. Mass.vC. formosa, Sarg. Allied to C. pruinosa. Shrub, to 15 ft.: Lvs. oblong-ovate, rounded or cuneate at the base, slightly lobed, slightly hairy above while young: corymbs many-fld.: fr. ovoid or obovoid, scarlet, pruinose, with 4-5 stones. N. Y.{{SCH}}

*C. geneseensis. Sarg. Allied to C. Crus-galli. Small tree, to 12 ft., spiny, glabrous: Lvs. obovate- oblong, pointed at the rounded or acute apex, with prominent veins: corymbs many-fld., lax; anthers pink: fr. ovoid, scarlet, ½ in. long, with 1-3 nutlets, in Oct. W. N. Y. G.C. III. 53:115.{{SCH}}

*C. gloriosa, Sarg. Allied to C. pedicellata. Tree, to 25 ft., with few spines: Lvs. ovate, cuneate or rounded at the base, rough above, slightly pubescent on the veins below, sometimes finally glabrous: corymbs 10-15-fld.; stamens 7- 10: fr. ovoid, often unsymmetrical, deep crimson, in Sept. W. N. Y. {{SCH}}

*C. grandiflora, Koch (C. lobata, Bosc. Cratae-mespilus grandiflora, Camus). Small tree: Lvs. elliptic, serrate, often slightly lobed toward the apex, pubescent: fls. 1-3, large: fr. brown, globose, large. Supposed to be a hybrid between Mespilus germanica and a Crataegus. G.F. 10:35. R.H. 1869, p. 80.{{SCH}}

*C. Harbisonii. Beadle. Belongs to group Bracteatae allied to Intricate. Tree, to 25 ft.: Lvs. oval or broadly obovate, coarsely serrate; petioles glandular: corymbs many-fld., with conspicuous glandular bracts: fr. red or bright red, in Oct. S.S. 13:691.{{SCH}}

*C. heterophylla, Fluegge. Allied to C. monogyna. Lvs. larger, usually trifid: fr. larger, bright red: corymbs many-fld. B.R. 14:1161:22:1847.{{SCH}}

*C. Holmesiana, Sarg. Allied to C. pedicellata. Tree, to 30 ft.: Lvs. oval or ovate, slightly lobed, at maturity yellowish green, glabrous: fls. ½-¾ in. across; stamens usually 5, anthers purple: fr. ovoid, crimson, with usually 3 stones. Montreal to Pa., W. N. Y. and Ont. S.S. 13:676.{{SCH}}

* C. integriloba, Sarg. Allied to C. tomentosa. Tree, to 10 ft., spiny: Lvs. broadly obovate or oval, broadly cuneate at the base, slightly lobed. glabrous: corymbs villous; calyx-lobes entire: fr. subglobose, ⅓-½in- across, scarlet, lustrous. Que. G.C. III. 47:60.{{SCH}}

*C. irrosa, Sarg. Allied to C. pedicellata. Shrub, to 12 ft.: Lvs. ovate, cuneate or rounded at the base, slightly lobed, lustrous and glabrous above: stamens 20, anthers yellow: fr. ovoid, dark red, lustrous. Que.{{SCH}}

*C. Korolkowii, Henry =C. Wattiana. See also No. 50.{{SCH}}

*C. lauta, Sarg. Allied to C. Ellwangeriana. Arborescent shrub, spiny: Lvs. ovate, acuminate, scabrate above, sparingly pubescent on the veins below: corymbs 8-12-fld., compact: fr. ovoid, bright orange-red. ¾ in. long, with 5 nutlets, in Sept. Origin unknown, much planted in Boston parks.{{SCH}}

*C. leiophylla, Sarg. Allied to C. pruinosa. Slender intricate, spiny shrub, to 15 ft., glabrous: Lvs. broadly ovate, usually rounded or truncate at the base, dark dull blue-green above: corymbs 5-7- fld., compact; anthers yellow: fr. obovoid. bright red. ,½ in. long, with usually 4 stones, in Nov. W. N. Y.{{SCH}}

*C. livoniana, Sarg. Allied to C. Crus-galli. Tree, to 20 ft., spiny, glabrous: Lvs. oblong-obovate, acute or rounded at the apex, finely and often doubly serrate: corymbs lax, 10-18-fld.; calyx-lobes glandular-serrate: fr. subglobose to ovoid, dark crimson, ½ in- long, with 2-4 stones, in Oct. N. Y. S.T.S. 2:129.{{SCH}}

*C. maineana, Sarg. (C. leiophylla var. maineana, Eggleston). Allied to C. pruinosa. Tree-like shrub, to 15 ft., spiny, glabrous: lvs. ovate to deltoid, acuminate, hairy while young: corymbs many-fld.; stamens 10; anthers dark purple : fr. globose, scarlet, scarcely pruinose, about ½ in. thick , in. Oct. W.N.Y.{{SCH}}

*C. matura. Sarg. Allied to C. pastorum. Shrub, to 10 ft., with few spines, glabrous: Lvs. oval to ovate-oblong, usually cuneate at the base, dark green above, yellow-green below: corymbs many-fld.; stamens 5-10; anthers red: fr. ovoid, dark purplish crimson, ¾ in. long, in Aug. Vt., Mass., N. Y.{{SCH}}

*C. Maximowicxii. Schneid. (C. sanguinea var. villosa, Maxim.). -Allied to C. sanguinea. Small tree, to 20 ft: Lvs. ovate, slightly lobed. pilose below: corymbs densely pilose: fr. pilose when young, finally glabrous. Amurland, Manchuria.{{SCH}}

*C. Palmeri, Sarg. Allied to C. Crus-galli. Tree, to 25 ft.: Lvs. broadly ovate to oblong, rounded or acute at the apex, coarsely serrate, glabrous: corymbs glabrous; stamens 10, with yellow anthers: fr. dull green, tinged with red, in Oct. S.M. 381.{{SCH}}

*C. peregrina, Sarg. Allied to C. mollis. Tree: Lvs. ovate, broadly cuneate, with 5-6 pairs of narrow lobes, glabrous above, villous beneath: corymbs many-6d., villous: fr. ovate-globose, dark dull purple, pubescent at the base and apex. ½ in. across. Origin unknown, probably S. W. Asia. S.T.S. 2:191.{{SCH}}

*C. perjucunda, Sarg. Allied to C. pruinosa. Spiny shrub, glabrous: Lvs. ovate, acuminate, dark green above: corymbs 8-10-fld.; anthers white: fr. ovoid, orange-red, finally crimson, slender-stalked, less than ½ in. long, in Oct. Ont., N. Y.{{SCH}}

*C. praecox, Sarg. (C. praecoqua, Sarg.). Allied to C. rotundifolia. Shrub, to 10 ft., spiny: Lvs. rhomboidal to oval, slightly hairy while young, glabrous at maturity and scabrous above: corymbs slightly villous, many-fld.; stamens 10: fr. subglobose, dark crimson, ⅔-in. thick, in Aug. Vt., Que.{{SCH}}

*C. promisea, Sarg. Allied to C. pruinosa. Shrub, to 12 ft., spiny, glabrous: Lvs. oblong-ovate, acuminate, deeply lobed: corymbs lax, many-fid.; stamens 5-7; anthers pink: fr. ovoid, crimson, not pruinose, less than ½ in- long, in Sept. W. N. Y.{{SCH}}

*C. rivuaris, Nutt. Allied to C. Douglasii. Shrub: Lvs. ovate-lanceolate, serrate, glabrous at length. Wyo. to Colo, and Utah. 8.3.4:176.{{SCH}}

*C. Sargentii. Beadle. Allied to C. intricata. Tree, to 20 ft.: Lvs. elliptic to oblong-ovate, slightly lobed, glabrous at maturity: corymbs slightly villous or glabrous; stamens 20, with purple anthers: fr. yellow or orange-yellow, tinged with red, in Sept. Ga. to Tenn. and Ala.{{SCH}}

*C. songarica, Regel= C. Wattiana.{{SCH}}

*C. spalkulata, Michx. Shrub or tree, to 20 ft.: Lvs. cuneate, oblanceolate, crenately serrate or 3-lobed at the apex: corymbs many-fld.: fr. scarlet, globular, ¼ in. across. Southern states. S.S. 4:185. B.R.22:1846 (as C. microcarpa). The only species of the group Microcarpae allied to the Apiifoliae.{{SCH}}

*C. triflora, Chapm. Shrub or small tree, to 20 ft.: lvs. ovate or elliptic, serrate, often slightly lobed, pubescent, 1-2 ½ in. long: corymbs 3-fld., hirsute; fls. 1 in. across; stamens 20; anthers yellow: fr. globose, red. Ga., Ala. Belongs to the group Triflorae, allied to Intricate.--Very distinct and handsome; has proved hardy at the Arnold Arboretum.{{SCH}}

*C. Wattiana, Hemsl. & Lace. (C. altaica, Lange. C. songarica, Regel). Allied to C. sanguinea. Lvs. smaller, truncate at the base, glabrous: corymbs glabrous: fr. yellow or reddish yellow, smaller. Cent. Asia. Var. incisa, Schneid. (C. Korolkowii, Henry. C. sanguinea var. incisa, Regel). Lvs. more1 deeply and acutely lobed. R.H. 1901:301.{{SCH}}

*C. Wilsonii, Sarg. Allied to C. tomentosa. Shrub, to 20 ft.: Lvs. ovate or obovate, acute or obtuse, lustrous above, sparingly villous beneath: fr. ovoid, red, nearly ½in. long, with 1-3 stones. Cent. China{{SCH}}

<gallery>

<gallery>

Image:Crataegus-monogyna-frugt.JPG|The fruit of Common Hawthorn (''C. monogyna'')

Image:Crataegus-monogyna-frugt.JPG|The fruit of Common Hawthorn (''C. monogyna'')

</gallery>

</gallery>

==References and external links ==

==References==

<references/>

<!--- xxxxx  *Flora: The Gardener's Bible, by Sean Hogan. Global Book Publishing, 2003. ISBN 0881925381  -->

* [http://www.bjucmp.edu.cn/zcy/Bh3-6.htm Photos of Chinese haw berries]

<!--- xxxxx  *American Horticultural Society: A-Z Encyclopedia of Garden Plants, by Christopher Brickell, Judith D. Zuk. 1996. ISBN 0789419432  -->

* [http://groups-beta.google.com/group/bionet.plants/msg/87b491ab63919f48?fwc=1 Key to Crataegus of Western North America]

<!--- xxxxx  *Sunset National Garden Book. Sunset Books, Inc., 1997. ISBN 0376038608  -->

* [http://www.armeniapedia.org/index.php?title=Crataegus Medicinal uses of Hawthorn in Armenia]

 

* [http://www.pfaf.org/leaflets/crataegs.php Edible uses of hawthorns at Plants For A Future]

==External links==

* [http://www.cas.vanderbilt.edu/bioimages/species/frame/crph.htm ''Crataegus phaenopyrum'' images at bioimages.vanderbilt.edu]

*{{wplink}}

 

[[Category:Categorize]]

[[Category:Maloideae]]

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