Difference between revisions of "Acacia"

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{{Taxobox
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{{SPlantbox
| color = lightgreen
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|familia=Fabaceae
| name = ''Acacia''
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|genus=Acacia
| image = Acacia melanoxylon branch.jpg
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|common_name=Acacia, thorntree, wattle
| image_width = 250px
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|lifespan=perennial
| image_caption = ''[[Acacia melanoxylon]]'' foliage and flowers
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|features=evergreen, deciduous
| regnum = [[Plant]]ae
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|Temp Metric=°F
| divisio = [[Flowering plant|Magnoliophyta]]
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|image=Acacia pycnantha Golden Wattle.jpg
| classis = [[Magnoliopsida]]
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|image_width=240
| ordo = [[Fabales]]
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|image_caption=Acacia pycnantha (Golden Wattle)
| familia = [[Fabaceae]]
 
| subfamilia = [[Mimosoideae]]
 
| tribus = Acacieae
 
| genus = '''''Acacia'''''
 
| genus_authority = [[Gerrit Smith Miller|Miller]]
 
| subdivision_ranks = Species
 
| subdivision =
 
About 1,300; see [[List of Acacia species]]
 
 
}}
 
}}
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'''''Acacia''''' ({{pron-en|əˈkeɪʃə}}) is a [[genus]] of [[shrub]]s and [[tree]]s belonging to the family [[Fabaceae]].  Acacias are also known as '''thorntrees''', '''[[whistling thorn]]s''' or '''wattles''', including the '''yellow-fever acacia''' and '''umbrella acacias'''.
  
[[Image:Acacia-tree.jpg|thumb|right|245px|Acacia tree in the [[Serengeti]], [[Tanzania]]]]
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Until 2005, there were thought to be roughly 1300 [[species]] of acacia worldwide, about 960 of them native to [[Australia]], with the remainder spread around the tropical to warm-[[temperate]] regions of both hemispheres, including [[Europe]], [[Africa]], southern [[Asia]], and the [[Americas]]. However, the genus was then divided into five, with the name ''Acacia'' retained for the Australian species, and most of the species outside Australia divided into ''[[Vachellia]]'' and ''[[Senegalia]].''
 
 
'''''Acacia''''' is a genus of [[shrub]]s and [[tree]]s of [[Gondwana|Gondwanian]] origin belonging to the subfamily [[Mimosoideae]] of the family [[Fabaceae]], first described from Africa by [[Carolus Linnaeus|Linnaeus]] in [[1773]].  
 
  
 
Acacias are also known as '''thorntrees''' or '''wattles''', including the yellow-fever acacia and umbrella acacias.   
 
Acacias are also known as '''thorntrees''' or '''wattles''', including the yellow-fever acacia and umbrella acacias.   
  
There are roughly 1300 species of ''Acacia'' worldwide, about 950 of them native to [[Australia]], with the remainder spread around the dry tropical to warm-temperate regions of both hemispheres, including Africa, southern Asia, and the Americas.  
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A few species are widely grown as ornamentals in [[garden]]s; the most popular perhaps is ''Acacia dealbata'' (Silver Wattle), with its attractive glaucous to silvery leaves and bright yellow flowers; it is erroneously known as "mimosa" in some areas where it is cultivated, through confusion with the related genus ''[[Mimosa]]''.
  
The genus ''Acacia'' however is apparently not [[monophyletic]]. This discovery has led to the breaking up of ''Acacia'' into five new genera as discussed in [[list of Acacia species]].
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The [[Leaf|leaves]] of acacias are compound [[pinnate]] in general. In some species, however, more especially in the Australian and [[Pacific islands]] species, the leaflets are suppressed, and the leaf-stalks ([[Petiole (botany)|petiole]]s) become vertically flattened, and serve the purpose of leaves. These are known as [[phyllode]]s. The vertical orientation of the phyllodes protects them from intense sunlight, as with their edges towards the sky and earth they do not intercept light so fully as horizontally placed leaves. A few species (such as ''[[Acacia glaucoptera]]'') lack leaves or phyllodes altogether, but possess instead [[cladode]]s, modified leaf-like [[photosynthesis|photosynthetic]] stems functioning as leaves.
  
==General discussion==
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The small [[flower]]s have five very small petals, almost hidden by the long [[stamens]], and are arranged in dense globular or cylindrical clusters; they are yellow or cream-colored in most species, whitish in some, even purple (''Acacia purpureapetala'') or red (''[[Acacia leprosa]]'' Scarlet Blaze). ''Acacia'' flowers can be distinguished from those of a large related genus, ''[[Albizia]]'', by their stamens which are not joined at the base. Also, unlike individual ''[[Mimosa]]'' flowers, those of ''Acacia'' have more than 10 stamens..<ref>{{cite book|last=Singh|first=Gurcharan|title=Plant Systematics: An Integrated Approach|publisher=Science Publishers|year=2004|pages=445|isbn=1578083516|url=http://books.google.com/books?id=In_Lv8iMt24C}}</ref>
The northernmost species in the genus is ''Acacia greggii'' ([[Catclaw Acacia]]), reaching 37°10' N in southern [[Utah]] in the [[United States]]; the southernmost are ''Acacia dealbata'' ([[Silver Wattle]]), ''Acacia longifolia'' ([[Coast Wattle]] or Sydney Golden Wattle), ''Acacia mearnsii'' ([[Black Wattle]]), and ''Acacia melanoxylon'' ([[Acacia melanoxylon|Blackwood]]), reaching 43°30' S in [[Tasmania]], Australia, while ''[[Acacia caven]]'' ([[Mimosa caven|Espinillo Negro]]) reaches nearly as far south in northeastern [[Chubut Province]] of [[Argentina]]. Australian species are usually called '''wattles''', while African and American species tend to be known as '''acacias'''.
 
  
[[Image:Koeh-003.jpg|thumb|left|''Acacia catechu'']]
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The plants often bear spines, especially those species growing in arid regions. These sometimes represent branches which have become short, hard and pungent, or sometimes leaf-[[stipule]]s. ''[[Acacia armata]]'' is the Kangaroo-thorn of Australia and ''[[Acacia erioloba]]'' is the Camelthorn of Africa.
The leaves of acacias are compound pinnate in general. In some species, however, more especially in the Australian and Pacific islands species, the leaflets are suppressed, and the leaf-stalks ('''[[Petiole (botany)|petiole]]s''') become vertically flattened, and serve the purpose of leaves; these are known as '''[[phyllode]]s'''. The vertical orientation of the phyllodes protects them from intense sunlight, as with their edges towards the sky and earth they do not intercept light so fully as horizontally placed leaves. A few species (such as ''[[Acacia glaucoptera]]'') lack leaves or phyllodes altogether, but possess instead '''[[cladode]]s''', modified leaf-like photosynthetic stems functioning as leaves.
 
  
The small [[flower]]s have five very small petals, almost hidden by the long stamens, and are arranged in dense globular or cylindrical clusters; they are yellow or cream-colored in most species, whitish in some, even purple (as in ''[[Acacia purpureapetala]]'') or red (in the recently grown cultivar ''[[Acacia leprosa]] 'Scarlet Blaze''').
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{{Inc|
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Acacia (from word meaning a point or thorn, referring to the stipules often spinescent). Leguminosae, tribe Mimoseae. Trees or shrubs grown out-of-doors in warmer parts of the United States and some of the species as cool greenhouse plants for the showy yellow bloom.
  
The plants often bear spines, especially those species growing in arid regions. These sometimes represent branches which have become short, hard and pungent, or sometimes leaf-stipules. ''Acacia armata'' is the [[Kangaroo-thorn]] of Australia, ''Acacia giraffae'', the [[Acacia erioloba|Camelthorn]] of Africa. In the Central American ''[[Acacia sphaerocephala]]'', ''[[Acacia spadicigera]]'', ''[[Acacia cornigera]]'', and ''[[Acacia collinsii]]'' (collectively known as the bullthorn acacias), the large thorn-like stipules are hollow and afford shelter for [[ant]]s, which feed on a secretion of sap on the leaf-stalk and small, lipid-rich food-bodies at the tips of the leaflets called Beltian bodies; in return they usually protect the plant against [[herbivores]]. Some species of ants will also trim competing p
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Leaves bipinnate or reduced to phyllodia with vertical edges (i.e., lf.-like petioles): fls. regular, orange-yellow, occasionally lemon-yellow or white, in cylindrical spikes or globular heads, solitary, or in pairs or clusters, or in axillary racemes; sepals and petals 5, 4 or 3, free or united; stamens many, long: pod a legume, opening by two valves (occasionally indehiscent): funicle of the seed filiform or ending in club-shaped aril, either twice encircling the seed or simply bent back upon itself. (The difference between Acacia and Albizzia lies in the stamens, which are free in the former and united at base in the latter.) A very large genus (said to be 450 species) dispersed throughout the tropical parts of the earth and even pushing their way into parts of the temperate zones. The phyllodine series is confined almost entirely to Australia and the Pacific Isls., while the bipinnate series is scattered over the warm parts of the remainder of the globe. The number of species reported from Amer. is large (about 70), of which at least 30 are Mexican. Acacias are said to be natives of the following states: Ariz., Ark., Calif., Fla., Okla., Kan., La.. Mo., Nev., New Mex. and Texas. Those in the following account are Australian, unless otherwise stated. This list will undoubtedly be modified as botanists segregate the other genera from the Mimoseae group. Acacias vary greatly under cult., the variation affecting infl., size and shape of lf., and even the funicle. One should not expect to determine an unfamiliar species without lvs., fls. and fr., with its seeds in place.
lants around the acacia, while other ant species will do nothing to benefit their host.
 
  
In common parlance the term "acacia" is occasionally misapplied to species of the genus ''Robinia'', which also belongs in the pea family. ''[[Robinia pseudoacacia]]'', an American species locally known as Black locust, is sometimes called "false acacia" in cultivation in Britain.
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Other species interesting because of their ant-inhabited thorns are described under Bull-horn Acacias. Other species referred to Acacia are to be found under Albizzia.
  
In Australia, ''Acacia'' species are sometimes used as food plants by the [[larva]]e of [[Hepialidae|hepialid]] [[moth]]s of the genus ''[[Aenetus]]'' including ''A. ligniveren''. These burrow horizontally into the trunk then vertically down. Other [[Lepidoptera]] larvae which have been recorded feeding on ''Acacia'' include [[Brown-tail]], ''[[Endoclita|Endoclita malabaricus]]'' and [[Turnip Moth]]. The leaf-mining larvae of some [[Bucculatricidae|bucculatricid]] moths also feed on ''Acacia'': ''Bucculatrix agilis'' feeds exclusively on ''Acacia horrida'', ''Bucculatrix flexuosa'' feeds exclusively on ''Acacia nilotica''.
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Acacias are quick-growing plants and are short-lived. Various kinds have been known to grow from 11 to 12 feet in four months and 25 feet in six years. These trees are thus in their full maturity at thirty years of age, and shortly afterward begin to deteriorate. While they may thus be used as street trees to secure immediate effects, more permanent trees should also be planted to take their places. The leaves of some species are used in cookery, the flowers of A. Farnesiana for perfumery, the bark of various species for tanning; A. Senegal furnishes most of the gum arabic of commerce; a drug or medicine is made from the wood of A. Catechu; a soap or hair-wash from A. concinna; several are used as forage plants, others for dyes, and still others for fiber. Many of them have scented wood, others make fine furniture wood, and are used for cabinetwork and fence-posts or fuel, still others for street trees (as A. melanoxylon and A. dealbata), and all are more or less ornamental. The Ark of the Covenant, as well as the furniture of the Tabernacle, are said to have been made from timber of A. Seyal, which yields the Shittim wood of the Bible. Also on account of its incorruptible wood, this species for ages was used by the Egyptians to make coffins for the burial of their kings.
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{{SCH}}
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}}
  
== Uses ==
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==Cultivation==
===Industrial and medicinal uses===
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{{Inc|
Various species of acacia yield gum. True [[gum arabic]] is the product of ''[[Acacia Senegal]]'', abundant in dry tropical west Africa from [[Senegal]] to northern [[Nigeria]].  
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Acacias out-of-doors will not endure much frost. They seem to thrive very well in localities in which the winter temperature is as low as 20° F., or even 18°, and it is to be hoped that by careful selection strains may be secured that will withstand an even lower temperature. Some species are alkali-resistant as A. cyclops, A. retinodes and perhaps others. After the trees are once well established they do not ordinarily require further irrigation since they have a faculty of seeking water. However, if they are given plenty of water and good soil their growth is very rapid. Weevils sometimes ruin acacia seeds by laying their eggs in the flower-buds and appearing later in the pod. The cottony cushion scale and the black scale are also found to a limited extent, but so far have not proved troublesome.
  
''[[Acacia arabica]]'' is the gum-arabic tree of India, but yields a gum inferior to the true gum-arabic. The bark of ''Acacia arabica'', under the name of '''babul''' or '''babool''', is used in Scinde for tanning. In [[Ayurvedic medicine]], babul is considered a remedy that is helpful for treating premature ejaculation.
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Cultivation in greenhouses as florists' plants is confined to few species, perhaps not more than a dozen being commercially valuable. All of this most important section thrive in a winter temperature ranging from 40° to 50°; in fact, little above the freezing point is sufficient. They do not like heat, and consequently are not adapted for forcing. If wintered cool and allowed to come along naturally with the increasing heat and light of the spring, they will flower in March and April, a season when their graceful beauty is appreciated in the private conservatory or is valuable to the commercial florist. The prevailing color of all the Australian species is yellow, varying from pale lemon to deep orange. The tall-growing kinds, or rather those inclined to make long, straight shoots, make excellent subjects for planting permanently against a glass partition of a conservatory, or against a pillar. There is scarcely a more beautiful plant than A. pubescens, with its slightly drooping, yellow racemes. It deserves a favored place in every cool conservatory. The acacias are of easy culture. If planted permanently in the border, provision for drainage should be made. A good, coarse, turfy loam, of not too heavy texture, is all they want, with the addition of a fifth part of leaf-mold or well-rotted spent hops. Few of our greenhouse pests trouble them. Water in abundance they like at all times, and in their growing season, which is the early summer months, a daily syringing is necessary. Several of the species of bushy habit are very much grown as pot-plants in Europe, and are now largely imported and sold for the eastern trade. A. armata and A. Drummondii are good species for this purpose. With our hot summers, the commercial man will do better to import than to attempt to grow them from cuttings. The acacias need pruning, or they will soon grow straggling and unshapely; more especially is this true of those grown in pots. After flowering, cut back the leading shoots rather severely. Shift into a larger pot if roots demand it, and encourage growth by a genial heat and syringing, giving at same time abundance of light and air. They should be plunged out-of-doors as soon as danger of frost is past, and removed to the greenhouse before any danger of early fall frosts. Cuttings root surely but not quickly. The best material is the side shoots from a main stem in the condition that florists call half-ripened —that is, not green and succulent as for a verbena, nor as firm and hard as the wood of a hybrid perpetual rose in November. The wood or shoot will be in about the right condition in June. No bottom heat is needed, but the cuttings should be covered with a close frame and kept moderately moist and cool by shading. The following spring these young plants can be either planted out-of-doors, where there is a good chance to keep them well watered, or grown on in pots, as described above. A few of the finest species are A. pubescens, suitable for training on pillars; A. Riceana makes a bush or can be trained; A. longifolia, an erect species, deserves a permanent position in the greenhouse border. Of all the species best adapted for medium-sized, compact pot-plants, A. armata and A. Drummondii are the best. The former has small, simple, dark green leaves and globular, pure yellow flowers. A. Drummondii has drooping, cylindrical, pale lemon flowers. As both these flower in March without any forcing in our northern greenhouses, they are very valuable acquisitions to our Easter plants. The acacia has two distinctive charms: the foliage is either small, simple and glaucous, as in A. armata, or much divided, graceful and fern-like, as in A. pubescent. All the acacias are among the freest-flowering of our hard-wooded plants. {{SCH}}
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}}
  
The bark of various Australian species, known as wattles, is very rich in tannin and forms an important article of export; important species include ''Acacia pycnantha'' (Golden Wattle), ''Acacia decurrens'' (Tan Wattle), ''Acacia dealbata'' (Silver Wattle) and ''Acacia mearnsii'' (Black Wattle). Black Wattle is grown in  plantations in South Africa. Most Australian acacia species introduced to South Africa, have become an enormous problem due to their naturally aggressive propagation. The pods of ''[[Acacia nilotica]]'' (under the name of neb-neb), and of other African species are also rich in tannin and used by tanners.
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===Propagation===
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Acacia seeds can be difficult to germinate. Research has found that immersing the seeds in various temperatures (usually around 80 °C) and manual seed coat chipping can improve yields to approximately 80 percent.<ref>{{cite journal |author=J Clemens, PG Jones, NH Gilbert|date=|year=1977|month=|title=Effect of seed treatments on germination in Acacia|journal=Australian Journal of Botany |volume=25 |issue=3 |pages=269-267 |pmid=|doi=10.1071/BT9770269|url=http://www.publish.csiro.au/nid/65/paper/BT9770269.htm}}</ref>
  
[[Image:Australian_Golden_Wattle_Blossums.jpg|thumb|Australian Golden Wattle flowers]]
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{{Inc|
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Propagation is by seeds sown either under glass or out in the open ground, or by cuttings from half-ripened wood, taken with a heel. Seeds may be prepared for planting in two ways: First, place them in ashes among the dying embers of a fire and allow them to remain until cool. These do not require immediate sowing, but if they are sown they will not perish if rain does not fall very soon afterwards. Second, pour hot water over seed, let cool and soak from twelve to forty-eight hours. Sow without allowing seeds to become dry. Either method softens the hard seed-coats and hastens germination. They will then usually germinate in about seven days to three or four weeks, depending upon the species and the season in which they are sown. Seed may be sown in the propagating-house at any time throughout the year, though early spring is the natural time. For open ground, sow in March or April. After germination, the plants are pricked off into flats or pots and shifted into larger ones as occasion requires. They are thus kept in pots until they are ready to be transplanted to their permanent quarters, since if placed in the open ground at once the tap-roots will grow with too great rapidity and the tree will either have to be balled or transplanted with the greatest care to prevent its receiving a shock, from which it will take at least a year to recover. When buying seedlings from a nursery, therefore, reject all those whose roots have penetrated the pot. While several species (A. pycnantha, A. melanoxylon, A. decurrens var. dealbata, etc.) have been known to resow themselves in California, there is no danger of their becoming a pest (such as A. armata in Australia), since the seedlings are seldom able to live through the dry season without irrigation. Cuttings should be made from the half-ripened wood, of which the best are from the side shoots of the main stem, taken with a heel. No bottom heat is required, or very little, but they should be covered with a light frame and kept moist and cool by shading. They root slowly but freely and should be potted immediately after rooting, but should not be planted in the open soil until they have developed good roots.
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{{SCH}}
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}}
  
Most acacia species are used for valuable timber; such are ''Acacia melanoxylon'' (Blackwood) from [[Australia]], which attains a great size; its wood is used for furniture, and takes a high polish; and ''Acacia homalophylla'' ([[Myall Wood]], also Australian), which yields a fragrant timber, used for ornamental purposes. ''[[Acacia formosa]]'' supplies the valuable [[Cuba]]n timber called sabicu. ''[[Acacia seyal]]'' is thought to be the shittah tree of the [[Bible]], which supplied shittim-wood. This was used in the construction of the [[Ark of the Covenant]]. ''[[Acacia heterophylla]]'' from [[Réunion]] island, and ''[[Koa|Acacia koa]]'' from the [[Hawaiian Islands]] are excellent timber trees.
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===Pests and diseases===
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In Australia, ''Acacia'' species are sometimes used as food plants by the [[larva]]e of [[Hepialidae|hepialid]] [[moth]]s of the [[genus]] ''[[Aenetus]]'' including ''A. ligniveren''. These burrow horizontally into the trunk then vertically down. Other [[Lepidoptera]] larvae which have been recorded feeding on ''Acacia'' include [[Brown-tail]], ''[[Endoclita|Endoclita malabaricus]]'' and [[Turnip Moth]]. The leaf-mining larvae of some [[Bucculatricidae|bucculatricid]] [[moth]]s also feed on ''Acacia'': ''[[idae|Bucculatrix agilis]]'' feeds exclusively on ''[[Acacia horrida]]'' and ''[[Bucculatricidae|Bucculatrix flexuosa]]'' feeds exclusively on ''[[Acacia nilotica]]''.
  
''Acacia farnesiana'' is used in the perfume industry due to its strong fragrance.
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== Species ==
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There are over 1,300 species of Acacia. See [[List of Acacia species]] for a complete listing.  The genus however is apparently not [[monophyletic]]. This discovery has led to the breaking up of ''Acacia'' into five new genera as discussed in [[list of Acacia species]].
  
An astringent medicine, called [[catechu]] or cutch, is procured from several species, but more especially from ''[[Acacia catechu]]'', by boiling down the wood and evaporating the solution so as to get an extract.
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{{#ask: [[genus::Acacia]]
 
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| ?common_name
===Ornamental uses===
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| ?habit
[[Image:Acaciaauriculiformis1web.jpg|thumb|right|Earpod Wattle (''Acacia auriculiformis'')]]
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| ?lifespan
A few species are widely grown as ornamentals in [[garden]]s; the most popular perhaps is ''Acacia dealbata'' (Silver Wattle), with its attractive glaucous to silvery leaves and bright yellow flowers; it is erroneously known as "mimosa" in some areas where it is cultivated, through confusion with the related genus ''[[Mimosa]]''.
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| ?exposure
 
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| ?water
===Culinary uses===
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| ?min_temp
[[Acacia seed]]s are often used for food and a variety of other products. The seeds of ''[[Acacia niopo]]'', for instance, are roasted and used as [[snuff]] in [[South America]].
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}}
 
 
In [[Myanmar|Burma]], [[Laos]] and [[Thailand]], the feathery shoots of ''[[Acacia pennata]]'' (common name ''cha-om'', ชะอม) are used in soups, curries, omelette
 
s, and stir-fries.
 
 
 
Honey made from the acacia flower is considered a delicacy, appreaciated for its mild flowery taste, soft running texture and glass like appearance.  Acacia nectar can be an abundant enough to classify the honey as [[monofloral honey]].
 
 
 
===Pharmacological uses===
 
Many Acacia species contain some psychoactive alkaloids of which [[Dimethyltryptamine|DMT]] and [[N-methyltryptamine|NMT]] are the most prominent and useful. The leaves, stems and/or roots can be made into a brew together with some [[Monoamine oxidase inhibitor|MAOI]]-containing plant to obtain an effect when taken orally. This could be seen as a kind of ''[[Ayahuasca]]''. Maybe in relation to this effect, [[Egyptian mythology]] has associated the acacia tree with characteristics of the [[tree of life]] (cf. article on the [[Legend of Osiris and Isis]]).  As a spiritual icon it is also one of the most powerful symbols in [[freemasonry]], representing the eternal soul and purity of the soul.
 
 
 
Alkaloids in different species, from ''[[TiHKAL]]'' (by [[Alexander Shulgin]]):
 
(There remains some question over whether any or all specimens of the following species would contain the alkaloids reported in TIHKAL, but it's a starting point. K Trouts books and notes on Acacia and tryptamines contain more recent information)
 
 
 
{|
 
|-----
 
| ''[[Cootamundra wattle|Acacia baileyana]]''
 
| 0.02% tryptamine and β-carbolines, in the leaf
 
|-----
 
| ''[[Acacia maidenii]]'' || DMT and NMT, in the stem bark
 
|-----
 
| ''A. albida'' || DMT, in the leaf
 
|-----
 
| ''A. confusa''
 
| DMT and NMT, in the leaf, stem and bark
 
|-----
 
| ''[[Acacia cultriformis]]'' || tryptamine, in the leaf and stem
 
|-----
 
| ''A. laeta'' || DMT, in the leaf
 
|-----
 
| ''[http://plants.usda.gov/java/profile?symbol=ACME8 A. mellifera]''
 
| DMT, in the leaf
 
|-----
 
| ''[[Acacia melanoxylon]]'' || DMT, in the bark and leaf
 
|-----
 
| ''[[Acacia nilotica]]'' || DMT, in the leaf
 
|-----
 
| ''A. phlebophylla'' || DMT, in the leaf
 
|-----
 
| ''A. podalyriaefolia'' || tryptamine, in the leaf
 
|-----
 
| ''A. Senegal'' || DMT, in the leaf
 
|-----
 
| ''A. seyal'' || DMT, in the leaf
 
|-----
 
| ''A. sieberiana'' || DMT, in the leaf)
 
|-----
 
| ''A. simplicifolia''
 
| DMT and NMT, in the leaf, stem and trunk bark
 
|-----
 
| ''A. vestita'' || tryptamine, in the leaf and stem
 
|}
 
  
Acacia is also a very good and healthy soluble fiber, often used by [[IBS]] sufferers {{Fact|date=February 2007}}.
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==Gallery==
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<gallery perrow=5>
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File:Acacia dealbata.jpg|''[[Acacia dealbata]]''
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Image:Koeh-003.jpg|''Acacia catechu''
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Image:Acacia pendula-weeping acacia-IMG 1084 oc.jpg|Acacia pendula (weeping acacia)
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Image:Acacia_craspedocarpa-broad-leaved_mulga-IMG_0904_hunt07.jpg|Acacia craspedocarpa (broad-leaved mulga)
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Image:Acacia_craspedocarpa-broad-leaved_mulga-IMG_0905_hunt07.jpg|Acacia craspedocarpa (broad-leaved mulga)
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Image:Acacia_decurrens-var._moilis-IMG_2929_lokrun.jpg|Acacia decurrens var. moilis
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Image:Acacia_decurrens-var._moilis-IMG_2930_lokrun.jpg|Acacia decurrens var. moilis
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Image:Acacia_farnesiana-IMG_2911_lokrun.jpg|Acacia farnesiana
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Image:Acacia_farnesiana-IMG_2912_lokrun.jpg|Acacia farnesiana
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Image:Acacia_glaucoptera-flat_wattle-IMG_0910_hunt07.jpg|Acacia glaucoptera (flat wattle)
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Image:Acacia_glaucoptera-flat_wattle-IMG_0911_hunt07.jpg|Acacia glaucoptera (flat wattle)
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Image:Acacia_longifolia-IMG_2926_lokrun.jpg|Acacia longifolia
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Image:Acacia_longifolia-IMG_2927_lokrun.jpg|Acacia longifolia
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File:Acacia_greggii_thorns.jpg|Acacia greggii thorns
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</gallery>
  
== Species ==
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==References==
There are over 1,300 species of Acacia. See [[List of Acacia species]] for a complete listing.  
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<!--- xxxxx  *Flora: The Gardener's Bible, by Sean Hogan. Global Book Publishing, 2003. ISBN 0881925381  -->
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<!--- xxxxx  *American Horticultural Society: A-Z Encyclopedia of Garden Plants, by Christopher Brickell, Judith D. Zuk. 1996. ISBN 0789419432  -->
 +
<!--- xxxxx  *Sunset National Garden Book. Sunset Books, Inc., 1997. ISBN 0376038608  -->
  
== External links ==
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==External links==
{{wikispecies|Acacia}}
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*{{wplink}}
{{Commons|Acacia}}
 
* [http://www.worldwidewattle.com/ World Wide Wattle]
 
* [http://waynesword.palomar.edu/plaug99.htm Wayne's Word] on "The Unforgettable Acacias"
 
* [http://mulga.yage.net/acacia/ The genus Acacia and Entheogenic Tryptamines, with reference to Australian and related species, by mulga]
 
  
[[Category:Acacia| ]]
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__NOTOC__
[[Category:Pantropical flora]]
 
[[Category:Flora of Australia]]
 
[[Category:Flora of Argentina]]
 
[[Category:Trees of Africa]]
 

Latest revision as of 18:01, 23 August 2015


Acacia pycnantha (Golden Wattle)


Plant Characteristics
Lifespan: perennial
Cultivation
Features: evergreen, deciduous
Scientific Names

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Acacia >


Acacia (pronounced /əˈkeɪʃə/) is a genus of shrubs and trees belonging to the family Fabaceae. Acacias are also known as thorntrees, whistling thorns or wattles, including the yellow-fever acacia and umbrella acacias.

Until 2005, there were thought to be roughly 1300 species of acacia worldwide, about 960 of them native to Australia, with the remainder spread around the tropical to warm-temperate regions of both hemispheres, including Europe, Africa, southern Asia, and the Americas. However, the genus was then divided into five, with the name Acacia retained for the Australian species, and most of the species outside Australia divided into Vachellia and Senegalia.

Acacias are also known as thorntrees or wattles, including the yellow-fever acacia and umbrella acacias.

A few species are widely grown as ornamentals in gardens; the most popular perhaps is Acacia dealbata (Silver Wattle), with its attractive glaucous to silvery leaves and bright yellow flowers; it is erroneously known as "mimosa" in some areas where it is cultivated, through confusion with the related genus Mimosa.

The leaves of acacias are compound pinnate in general. In some species, however, more especially in the Australian and Pacific islands species, the leaflets are suppressed, and the leaf-stalks (petioles) become vertically flattened, and serve the purpose of leaves. These are known as phyllodes. The vertical orientation of the phyllodes protects them from intense sunlight, as with their edges towards the sky and earth they do not intercept light so fully as horizontally placed leaves. A few species (such as Acacia glaucoptera) lack leaves or phyllodes altogether, but possess instead cladodes, modified leaf-like photosynthetic stems functioning as leaves.

The small flowers have five very small petals, almost hidden by the long stamens, and are arranged in dense globular or cylindrical clusters; they are yellow or cream-colored in most species, whitish in some, even purple (Acacia purpureapetala) or red (Acacia leprosa Scarlet Blaze). Acacia flowers can be distinguished from those of a large related genus, Albizia, by their stamens which are not joined at the base. Also, unlike individual Mimosa flowers, those of Acacia have more than 10 stamens..[1]

The plants often bear spines, especially those species growing in arid regions. These sometimes represent branches which have become short, hard and pungent, or sometimes leaf-stipules. Acacia armata is the Kangaroo-thorn of Australia and Acacia erioloba is the Camelthorn of Africa.


Read about Acacia in the Standard Cyclopedia of Horticulture 

Acacia (from word meaning a point or thorn, referring to the stipules often spinescent). Leguminosae, tribe Mimoseae. Trees or shrubs grown out-of-doors in warmer parts of the United States and some of the species as cool greenhouse plants for the showy yellow bloom.

Leaves bipinnate or reduced to phyllodia with vertical edges (i.e., lf.-like petioles): fls. regular, orange-yellow, occasionally lemon-yellow or white, in cylindrical spikes or globular heads, solitary, or in pairs or clusters, or in axillary racemes; sepals and petals 5, 4 or 3, free or united; stamens many, long: pod a legume, opening by two valves (occasionally indehiscent): funicle of the seed filiform or ending in club-shaped aril, either twice encircling the seed or simply bent back upon itself. (The difference between Acacia and Albizzia lies in the stamens, which are free in the former and united at base in the latter.) A very large genus (said to be 450 species) dispersed throughout the tropical parts of the earth and even pushing their way into parts of the temperate zones. The phyllodine series is confined almost entirely to Australia and the Pacific Isls., while the bipinnate series is scattered over the warm parts of the remainder of the globe. The number of species reported from Amer. is large (about 70), of which at least 30 are Mexican. Acacias are said to be natives of the following states: Ariz., Ark., Calif., Fla., Okla., Kan., La.. Mo., Nev., New Mex. and Texas. Those in the following account are Australian, unless otherwise stated. This list will undoubtedly be modified as botanists segregate the other genera from the Mimoseae group. Acacias vary greatly under cult., the variation affecting infl., size and shape of lf., and even the funicle. One should not expect to determine an unfamiliar species without lvs., fls. and fr., with its seeds in place.

Other species interesting because of their ant-inhabited thorns are described under Bull-horn Acacias. Other species referred to Acacia are to be found under Albizzia.

Acacias are quick-growing plants and are short-lived. Various kinds have been known to grow from 11 to 12 feet in four months and 25 feet in six years. These trees are thus in their full maturity at thirty years of age, and shortly afterward begin to deteriorate. While they may thus be used as street trees to secure immediate effects, more permanent trees should also be planted to take their places. The leaves of some species are used in cookery, the flowers of A. Farnesiana for perfumery, the bark of various species for tanning; A. Senegal furnishes most of the gum arabic of commerce; a drug or medicine is made from the wood of A. Catechu; a soap or hair-wash from A. concinna; several are used as forage plants, others for dyes, and still others for fiber. Many of them have scented wood, others make fine furniture wood, and are used for cabinetwork and fence-posts or fuel, still others for street trees (as A. melanoxylon and A. dealbata), and all are more or less ornamental. The Ark of the Covenant, as well as the furniture of the Tabernacle, are said to have been made from timber of A. Seyal, which yields the Shittim wood of the Bible. Also on account of its incorruptible wood, this species for ages was used by the Egyptians to make coffins for the burial of their kings. CH


The above text is from the Standard Cyclopedia of Horticulture. It may be out of date, but still contains valuable and interesting information which can be incorporated into the remainder of the article. Click on "Collapse" in the header to hide this text.


Cultivation


Read about Acacia in the Standard Cyclopedia of Horticulture 

Acacias out-of-doors will not endure much frost. They seem to thrive very well in localities in which the winter temperature is as low as 20° F., or even 18°, and it is to be hoped that by careful selection strains may be secured that will withstand an even lower temperature. Some species are alkali-resistant as A. cyclops, A. retinodes and perhaps others. After the trees are once well established they do not ordinarily require further irrigation since they have a faculty of seeking water. However, if they are given plenty of water and good soil their growth is very rapid. Weevils sometimes ruin acacia seeds by laying their eggs in the flower-buds and appearing later in the pod. The cottony cushion scale and the black scale are also found to a limited extent, but so far have not proved troublesome.

Cultivation in greenhouses as florists' plants is confined to few species, perhaps not more than a dozen being commercially valuable. All of this most important section thrive in a winter temperature ranging from 40° to 50°; in fact, little above the freezing point is sufficient. They do not like heat, and consequently are not adapted for forcing. If wintered cool and allowed to come along naturally with the increasing heat and light of the spring, they will flower in March and April, a season when their graceful beauty is appreciated in the private conservatory or is valuable to the commercial florist. The prevailing color of all the Australian species is yellow, varying from pale lemon to deep orange. The tall-growing kinds, or rather those inclined to make long, straight shoots, make excellent subjects for planting permanently against a glass partition of a conservatory, or against a pillar. There is scarcely a more beautiful plant than A. pubescens, with its slightly drooping, yellow racemes. It deserves a favored place in every cool conservatory. The acacias are of easy culture. If planted permanently in the border, provision for drainage should be made. A good, coarse, turfy loam, of not too heavy texture, is all they want, with the addition of a fifth part of leaf-mold or well-rotted spent hops. Few of our greenhouse pests trouble them. Water in abundance they like at all times, and in their growing season, which is the early summer months, a daily syringing is necessary. Several of the species of bushy habit are very much grown as pot-plants in Europe, and are now largely imported and sold for the eastern trade. A. armata and A. Drummondii are good species for this purpose. With our hot summers, the commercial man will do better to import than to attempt to grow them from cuttings. The acacias need pruning, or they will soon grow straggling and unshapely; more especially is this true of those grown in pots. After flowering, cut back the leading shoots rather severely. Shift into a larger pot if roots demand it, and encourage growth by a genial heat and syringing, giving at same time abundance of light and air. They should be plunged out-of-doors as soon as danger of frost is past, and removed to the greenhouse before any danger of early fall frosts. Cuttings root surely but not quickly. The best material is the side shoots from a main stem in the condition that florists call half-ripened —that is, not green and succulent as for a verbena, nor as firm and hard as the wood of a hybrid perpetual rose in November. The wood or shoot will be in about the right condition in June. No bottom heat is needed, but the cuttings should be covered with a close frame and kept moderately moist and cool by shading. The following spring these young plants can be either planted out-of-doors, where there is a good chance to keep them well watered, or grown on in pots, as described above. A few of the finest species are A. pubescens, suitable for training on pillars; A. Riceana makes a bush or can be trained; A. longifolia, an erect species, deserves a permanent position in the greenhouse border. Of all the species best adapted for medium-sized, compact pot-plants, A. armata and A. Drummondii are the best. The former has small, simple, dark green leaves and globular, pure yellow flowers. A. Drummondii has drooping, cylindrical, pale lemon flowers. As both these flower in March without any forcing in our northern greenhouses, they are very valuable acquisitions to our Easter plants. The acacia has two distinctive charms: the foliage is either small, simple and glaucous, as in A. armata, or much divided, graceful and fern-like, as in A. pubescent. All the acacias are among the freest-flowering of our hard-wooded plants. CH


The above text is from the Standard Cyclopedia of Horticulture. It may be out of date, but still contains valuable and interesting information which can be incorporated into the remainder of the article. Click on "Collapse" in the header to hide this text.


Propagation

Acacia seeds can be difficult to germinate. Research has found that immersing the seeds in various temperatures (usually around 80 °C) and manual seed coat chipping can improve yields to approximately 80 percent.[2]


Read about Acacia in the Standard Cyclopedia of Horticulture 

Propagation is by seeds sown either under glass or out in the open ground, or by cuttings from half-ripened wood, taken with a heel. Seeds may be prepared for planting in two ways: First, place them in ashes among the dying embers of a fire and allow them to remain until cool. These do not require immediate sowing, but if they are sown they will not perish if rain does not fall very soon afterwards. Second, pour hot water over seed, let cool and soak from twelve to forty-eight hours. Sow without allowing seeds to become dry. Either method softens the hard seed-coats and hastens germination. They will then usually germinate in about seven days to three or four weeks, depending upon the species and the season in which they are sown. Seed may be sown in the propagating-house at any time throughout the year, though early spring is the natural time. For open ground, sow in March or April. After germination, the plants are pricked off into flats or pots and shifted into larger ones as occasion requires. They are thus kept in pots until they are ready to be transplanted to their permanent quarters, since if placed in the open ground at once the tap-roots will grow with too great rapidity and the tree will either have to be balled or transplanted with the greatest care to prevent its receiving a shock, from which it will take at least a year to recover. When buying seedlings from a nursery, therefore, reject all those whose roots have penetrated the pot. While several species (A. pycnantha, A. melanoxylon, A. decurrens var. dealbata, etc.) have been known to resow themselves in California, there is no danger of their becoming a pest (such as A. armata in Australia), since the seedlings are seldom able to live through the dry season without irrigation. Cuttings should be made from the half-ripened wood, of which the best are from the side shoots of the main stem, taken with a heel. No bottom heat is required, or very little, but they should be covered with a light frame and kept moist and cool by shading. They root slowly but freely and should be potted immediately after rooting, but should not be planted in the open soil until they have developed good roots. CH


The above text is from the Standard Cyclopedia of Horticulture. It may be out of date, but still contains valuable and interesting information which can be incorporated into the remainder of the article. Click on "Collapse" in the header to hide this text.


Pests and diseases

In Australia, Acacia species are sometimes used as food plants by the larvae of hepialid moths of the genus Aenetus including A. ligniveren. These burrow horizontally into the trunk then vertically down. Other Lepidoptera larvae which have been recorded feeding on Acacia include Brown-tail, Endoclita malabaricus and Turnip Moth. The leaf-mining larvae of some bucculatricid moths also feed on Acacia: Bucculatrix agilis feeds exclusively on Acacia horrida and Bucculatrix flexuosa feeds exclusively on Acacia nilotica.

Species

There are over 1,300 species of Acacia. See List of Acacia species for a complete listing. The genus however is apparently not monophyletic. This discovery has led to the breaking up of Acacia into five new genera as discussed in list of Acacia species.

 Common nameHabitLifespanExposureWaterMin temp
Acaciaperennial
Acacia acanthocladaperennialsun
Acacia acinaceaperennialsun
Acacia acuminata subsp. burkittiiperennialsun
Acacia aduncaperennialsun
Acacia aneuraperennialsun
Acacia aulacocarpaperennialsun
Acacia ausfeldiiperennialpart-sun
Acacia baileyanaperennialsun
Acacia berlandieriperennialsun
part-sun
Acacia binerviaperennialsun
Acacia boormaniiperennialsun
Acacia brachybotryaperennialsun
Acacia buxifoliaperennialsun
Acacia caffraperennialsun
Acacia calamifoliaperennialsun
Acacia cardiophyllaperennialsun
Acacia caveniaperennialsun
Acacia cognataperennialpart-sun
Acacia complanataperennialsun
Acacia covenyiperennialsun
Acacia crassaperennialsun
Acacia crassicarpaperennialsun
Acacia cultriformisperennialsun
Acacia cyclopsperennialsun
Acacia dealbataperennialsun
Acacia decoraperennialsun
Acacia decurrensperennialsun
Acacia drummondiiperennialsun
part-sun
Acacia dunniiperennialsun
Acacia elataperennialsun
Acacia estrophiolataperennialsun
Acacia fimbriataperennialsun
Acacia floribundaperennialpart-sun
Acacia giraffaeperennialsun
Acacia glaucopteraperennialpart-sun
Acacia greggiiperennial
Acacia hakeoidesperennialsun
Acacia harpophyllaperennialsun
Acacia havilandiorumperennialsun
Acacia howittiiperennialsun
Acacia implexaperennialsun
Acacia karrooperennialsun
Acacia lanigeraperennialsun
Acacia leptostachyaperennialsun
Acacia leucocladaperennialsun
Acacia longifoliaperennialsun
Acacia mangiumperennialsun
Acacia mearnsiiperennialsun
Acacia melanoxylonperennialsun
... further results

Gallery

References

External links


  1. Singh, Gurcharan (2004). Plant Systematics: An Integrated Approach. Science Publishers. pp. 445. ISBN 1578083516. http://books.google.com/books?id=In_Lv8iMt24C. 
  2. J Clemens, PG Jones, NH Gilbert (1977). "Effect of seed treatments on germination in Acacia". Australian Journal of Botany 25 (3): 269-267. doi:10.1071/BT9770269. http://www.publish.csiro.au/nid/65/paper/BT9770269.htm.